Overview of Homo sapiens Evolution
Overview of Homo sapiens Evolution
For most of their history, humans were nomadic hunter-gatherers. Subfamily: Homininae
Humans began exhibiting behavioral modernity about 160,000– Tribe: Hominini
60,000 years ago. The Neolithic Revolution, which began in
Genus: Homo
Southwest Asia around 13,000 years ago (and separately in a
few other places), saw the emergence of agriculture and Species: H. sapiens
permanent human settlement; in turn, this led to the development Binomial name
of civilization and kickstarted a period of continuous (and
Homo sapiens
ongoing) population growth and rapid technological change.
Linnaeus, 1758
Since then, a number of civilizations have risen and fallen, while
a number of sociocultural and technological developments have
resulted in significant changes to the human lifestyle.
Humans are omnivorous, capable of consuming a wide variety of plant and animal material, and have used
fire and other forms of heat to prepare and cook food since the time of Homo erectus. Humans can survive
for up to eight weeks without food and several days without water. Humans are generally diurnal, sleeping
on average seven to nine hours per day. Childbirth is dangerous, with a high risk of complications and
death. Often, both the mother and the father provide care for their children, who are helpless at birth.
Humans have a large, highly developed, and complex prefrontal cortex, the region of the brain associated
with higher cognition. Humans are highly intelligent and capable of episodic memory; they have flexible
facial expressions, self-awareness, and a theory of mind. The human mind is capable of introspection,
private thought, imagination, volition, and forming views on existence. This has allowed great technological
advancements and complex tool development through complex reasoning and the transmission of
knowledge to subsequent generations through language.
Evolution
Humans are apes (superfamily Hominoidea).[11] The lineage of apes that eventually gave rise to humans
first split from gibbons (family Hylobatidae) and orangutans (genus Pongo), then gorillas (genus Gorilla),
and finally, chimpanzees and bonobos (genus Pan). The last split, between the human and chimpanzee–
bonobo lineages, took place around 8–4 million years ago, in the late Miocene epoch.[12][13] During this
split, chromosome 2 was formed from the joining of two other chromosomes, leaving humans with only 23
pairs of chromosomes, compared to 24 for the other apes.[14] Following their split with chimpanzees and
bonobos, the hominins diversified into many species and at least two distinct genera. All but one of these
lineages – representing the genus Homo and its sole extant species Homo sapiens – are now extinct.[15]
The "out of Africa" migration took place in at least two waves, the first
around 130,000 to 100,000 years ago, the second (Southern Dispersal) around 70,000 to 50,000 years
ago.[27][28] H. sapiens proceeded to colonize all the continents and larger islands, arriving in Eurasia
125,000 years ago,[29][30] Australia around 65,000 years ago,[31] the Americas around 15,000 years ago,
and remote islands such as Hawaii, Easter Island, Madagascar, and New Zealand in the years 300 to 1280
CE.[32][33]
Human evolution was not a simple linear or branched progression but involved interbreeding between
related species.[34][35][36] Genomic research has shown that hybridization between substantially diverged
lineages was common in human evolution.[37] DNA evidence suggests that several genes of Neanderthal
origin are present among all non sub-Saharan-African populations, and Neanderthals and other hominins,
such as Denisovans, may have contributed up to 6% of their genome to present-day non sub-Saharan-
African humans.[34][38][39]
Human evolution is characterized by a number of morphological, developmental, physiological, and
behavioral changes that have taken place since the split between the last common ancestor of humans and
chimpanzees. The most significant of these adaptations are hairlessness,[40] obligate bipedalism, increased
brain size and decreased sexual dimorphism (neoteny). The relationship between all these changes is the
subject of ongoing debate.[41]
Hylobatidae (gibbons)
Pongo
Homininae (hominines)
P anina
Hominini (hominins)
History
Prehistory
Until about 12,000 years ago, all humans lived
as hunter-gatherers.[42][43] The Neolithic
Revolution (the invention of agriculture) first
took place in Southwest Asia and spread
through large parts of the Old World over the
following millennia.[44] It also occurred
independently in Mesoamerica (about 6,000
years ago),[45] China,[46][47] Papua New
Guinea,[48] and the Sahel and West Savanna Overview map of the peopling of the world by early human
regions of Africa.[49][50][51] migration during the Upper Paleolithic, following the
Southern Dispersal paradigm
Access to food surplus led to the formation of permanent human settlements, the domestication of animals
and the use of metal tools for the first time in history. Agriculture and sedentary lifestyle led to the
emergence of early civilizations.[52][53][54]
Ancient
An urban revolution took place in the 4th millennium BCE with the
development of city-states, particularly Sumerian cities located in
Mesopotamia.[55] It was in these cities that the earliest known form
of writing, cuneiform script, appeared around 3000 BCE.[56] Other
major civilizations to develop around this time were Ancient Egypt
and the Indus Valley Civilisation.[57] They eventually traded with
each other and invented technology such as wheels, plows and
sails.[58][59][60][61] Emerging by 3000 BCE, the Caral–Supe Great Pyramids of Giza, Egypt
civilization is the oldest complex civilization in the Americas.[62]
Astronomy and mathematics were also developed and the Great
Pyramid of Giza was built.[63][64][65] There is evidence of a severe drought lasting about a hundred years
that may have caused the decline of these civilizations,[66] with new ones appearing in the aftermath.
Babylonians came to dominate Mesopotamia while others,[67] such as the Poverty Point culture, Minoans
and the Shang dynasty, rose to prominence in new areas.[68][69][70] The Late Bronze Age collapse around
1200 BCE resulted in the disappearance of a number of civilizations and the beginning of the Greek Dark
Ages.[71][72] During this period iron started replacing bronze, leading to the Iron Age.[73]
In the 5th century BCE, history started being recorded as a discipline, which provided a much clearer
picture of life at the time.[74] Between the 8th and 6th century BCE, Europe entered the classical antiquity
age, a period when ancient Greece and ancient Rome flourished.[75][76] Around this time other civilizations
also came to prominence. The Maya civilization started to build cities and create complex calendars.[77][78]
In Africa, the Kingdom of Aksum overtook the declining Kingdom of Kush and facilitated trade between
India and the Mediterranean.[79] In West Asia, the Achaemenid Empire's system of centralized governance
became the precursor to many later empires,[80] while the Gupta Empire in India and the Han dynasty in
China have been described as golden ages in their respective regions.[81][82]
Medieval
Following the fall of the Western Roman Empire in 476, Europe entered the Middle Ages.[83] During this
period, Christianity and the Church would provide centralized authority and education.[84] In the Middle
East, Islam became the prominent religion and expanded into North Africa. It led to an Islamic Golden Age,
inspiring achievements in architecture, the revival of old advances in science and technology, and the
formation of a distinct way of life.[85][86] The Christian and Islamic worlds would eventually clash, with the
Kingdom of England, the Kingdom of France and the Holy Roman Empire declaring a series of holy wars
to regain control of the Holy Land from Muslims.[87]
In the Americas, between 200 and 900 CE Mesoamerica was in its Classic Period,[88] while further north,
complex Mississippian societies would arise starting around 800 CE.[89] The Mongol Empire would
conquer much of Eurasia in the 13th and 14th centuries.[90] Over this same time period, the Mali Empire in
Africa grew to be the largest empire on the continent, stretching from Senegambia to Ivory Coast.[91]
Oceania would see the rise of the Tuʻi Tonga Empire which expanded
across many islands in the South Pacific.[92] By the late 15th century, the
Aztecs and Inca had become the dominant power in Mesoamerica and the
Andes, respectively.[93]
Modern
The early modern period in Europe and the Near East (c. 1450–1800)
began with the final defeat of the Byzantine Empire, and the rise of the
Medieval French manuscript
Ottoman Empire.[94] Meanwhile, Japan entered the Edo period,[95] the
illustration of the three
Qing dynasty rose in China[96] and the Mughal Empire ruled much of classes of medieval society
India.[97] Europe underwent the Renaissance, starting in the 15th from the 13th-century Li
century,[98] and the Age of Discovery began with the exploring and Livres dou Santé
colonizing of new regions.[99] This included the the colonization of the
Americas[100] and the Columbian Exchange.[101] This expansion
led to the Atlantic slave trade[102] and the genocide of Native
American peoples.[103] This period also marked the Scientific
Revolution, with great advances in mathematics, mechanics,
astronomy and physiology.[104]
A tenuous balance of power among European nations collapsed in 1914 with the outbreak of the First
World War, one of the deadliest conflicts in history.[112] In the 1930s, a worldwide economic crisis led to
the rise of authoritarian regimes and a Second World War, involving almost all of the world's countries.[113]
The war's destruction led to the collapse of most global empires, leading to widespread decolonization.
Contemporary
Following the conclusion of the Second World War in 1945, the United States[114] and the USSR emerged
as the remaining global superpowers. This lead to a Cold War that saw a struggle for global influence,
including a nuclear arms race and a space race, ending in the collapse of the Soviet Union.[115][116] The
current Information Age, spurred by the development of the Internet and Artificial Intelligence systems, sees
the world becoming increasingly globalized and interconnected.[117]
Humans are one of the most adaptable species, despite having a low or narrow tolerance for many of the
earth's extreme environments.[125] Currently the species is present in all eight biogeographical realms,
although their presence in the Antarctic realm is very limited to research stations and annually there is a
population decline in the winter months of this realm. Humans established their nation-states in the other
seven realms, such as for example South Africa, India, Russia, Australia, Fiji, United States and Brazil
(each located in a different biogeographical realm).
By using advanced tools and clothing, humans have been able to extend their tolerance to a wide variety of
temperatures, humidities, and altitudes.[125][126] As a result, humans are a cosmopolitan species found in
almost all regions of the world, including tropical rainforest, arid desert, extremely cold arctic regions, and
heavily polluted cities; in comparison, most other species are confined to a few geographical areas by their
limited adaptability.[127] The human population is not, however, uniformly distributed on the Earth's
surface, because the population density varies from one region to another, and large stretches of surface are
almost completely uninhabited, like Antarctica and vast swathes of the ocean.[125][128] Most humans (61%)
live in Asia; the remainder live in the Americas (14%), Africa (14%), Europe (11%), and Oceania
(0.5%).[129]
Within the last century, humans have explored challenging environments such as Antarctica, the deep sea,
and outer space.[130] Human habitation within these hostile environments is restrictive and expensive,
typically limited in duration, and restricted to scientific, military, or industrial expeditions.[130] Humans have
briefly visited the Moon and made their presence felt on other celestial bodies through human-made robotic
spacecraft.[131][132][133] Since the early 20th century, there has been continuous human presence in
Antarctica through research stations and, since 2000, in space through habitation on the International Space
Station.[134]
In 2018, 4.2 billion humans (55%) lived in urban areas, up from 751 million in 1950.[144] The most
urbanized regions are Northern America (82%), Latin America (81%), Europe (74%) and Oceania (68%),
with Africa and Asia having nearly 90% of the world's 3.4 billion rural population.[144] Problems for
humans living in cities include various forms of pollution and crime,[145] especially in inner city and
suburban slums. Humans have had a dramatic effect on the environment. They are apex predators, being
rarely preyed upon by other species.[146] Human population growth, industrialization, land development,
overconsumption and combustion of fossil fuels have led to environmental destruction and pollution that
significantly contributes to the ongoing mass extinction of other forms of life.[147][148]
Biology
It is estimated that the worldwide average height for an adult human male is about 171 cm (5 ft 7 in), while
the worldwide average height for adult human females is about 159 cm (5 ft 3 in).[157] Shrinkage of stature
may begin in middle age in some individuals but tends to be typical in the extremely aged.[158] Throughout
history, human populations have universally become taller, probably as a consequence of better nutrition,
healthcare, and living conditions.[159] The average mass of an adult human is 59 kg (130 lb) for females
and 77 kg (170 lb) for males.[160][161] Like many other conditions, body weight and body type are
influenced by both genetic susceptibility and environment and varies greatly among individuals.[162][163]
Humans have a far faster and more accurate throw than other animals.[164] Humans are also among the best
long-distance runners in the animal kingdom, but slower over short distances.[165][153] Humans' thinner
body hair and more productive sweat glands help avoid heat exhaustion while running for long
distances.[166] Compared to other apes, the human heart produces greater stroke volume and cardiac output
and the aorta is proportionately larger.[167][168]
Genetics
Like most animals, humans are a diploid and eukaryotic species.
Each somatic cell has two sets of 23 chromosomes, each set
received from one parent; gametes have only one set of
chromosomes, which is a mixture of the two parental sets. Among
the 23 pairs of chromosomes, there are 22 pairs of autosomes and
one pair of sex chromosomes. Like other mammals, humans have
an XY sex-determination system, so that females have the sex
chromosomes XX and males have XY.[169] Genes and
environment influence human biological variation in visible
characteristics, physiology, disease susceptibility and mental
abilities. The exact influence of genes and environment on certain
traits is not well understood.[170][171]
A graphical representation of the
standard human karyotype, including
While no humans – not even monozygotic twins – are genetically
both the female (XX) and male (XY)
identical,[172] two humans on average will have a genetic similarity sex chromosomes.
of 99.5%-99.9%.[173][174] This makes them more homogeneous
than other great apes, including chimpanzees.[175][176] This small
variation in human DNA compared to many other species suggests a population bottleneck during the Late
Pleistocene (around 100,000 years ago), in which the human population was reduced to a small number of
breeding pairs.[177][178] The forces of natural selection have continued to operate on human populations,
with evidence that certain regions of the genome display directional selection in the past 15,000 years.[179]
The human genome was first sequenced in 2001[180] and by 2020 hundreds of thousands of genomes had
been sequenced.[181] In 2012 the International HapMap Project had compared the genomes of 1,184
individuals from 11 populations and identified 1.6 million single nucleotide polymorphisms.[182] African
populations harbor the highest number of private genetic variants. While many of the common variants
found in populations outside of Africa are also found on the African continent, there are still large numbers
that are private to these regions, especially Oceania and the Americas.[183] By 2010 estimates, humans have
approximately 22,000 genes.[184] By comparing mitochondrial DNA, which is inherited only from the
mother, geneticists have concluded that the last female common ancestor whose genetic marker is found in
all modern humans, the so-called mitochondrial Eve, must have lived around 90,000 to 200,000 years
ago.[185][186][187][188]
Life cycle
Most human reproduction takes place by internal fertilization via sexual intercourse, but can also occur
through assisted reproductive technology procedures.[189] The average gestation period is 38 weeks, but a
normal pregnancy can vary by up to 37 days.[190] Embryonic development in the human covers the first
eight weeks of development; at the beginning of the ninth week the embryo is termed a fetus.[191] Humans
are able to induce early labor or perform a caesarean section if the child needs to be born earlier for medical
reasons.[192] In developed countries, infants are typically 3–4 kg
(7–9 lb) in weight and 47–53 cm (19–21 in) in height at
birth.[193][194] However, low birth weight is common in developing
countries, and contributes to the high levels of infant mortality in
these regions.[195]
Both the mother and the father provide care for human offspring, in contrast to other primates, where
parental care is mostly done by the mother.[201] Helpless at birth, humans continue to grow for some years,
typically reaching sexual maturity at 15 to 17 years of age.[202][203][204] The human life span has been split
into various stages ranging from three to twelve. Common stages include infancy, childhood, adolescence,
adulthood and old age.[205] The lengths of these stages have varied across cultures and time periods but is
typified by an unusually rapid growth spurt during adolescence.[206] Human females undergo menopause
and become infertile at around the age of 50.[207] It has been proposed that menopause increases a woman's
overall reproductive success by allowing her to invest more time and resources in her existing offspring, and
in turn their children (the grandmother hypothesis), rather than by continuing to bear children into old
age.[208][209]
The life span of an individual depends on two major factors, genetics and lifestyle choices.[210] For various
reasons, including biological/genetic causes, women live on average about four years longer than men.[211]
As of 2018, the global average life expectancy at birth of a girl is estimated to be 74.9 years compared to
70.4 for a boy.[212][213] There are significant geographical variations in human life expectancy, mostly
correlated with economic development – for example, life expectancy at birth in Hong Kong is 87.6 years
for girls and 81.8 for boys, while in the Central African Republic, it is 55.0 years for girls and 50.6 for
boys.[214][215] The developed world is generally aging, with the median age around 40 years. In the
developing world, the median age is between 15 and 20 years. While one in five Europeans is 60 years of
age or older, only one in twenty Africans is 60 years of age or older.[216] In 2012, the United Nations
estimated that there were 316,600 living centenarians (humans of age 100 or older) worldwide.[217]
Human life stages
Diet
Humans are omnivorous, capable of consuming a wide variety of
plant and animal material.[218][219] Human groups have adopted a
range of diets from purely vegan to primarily carnivorous. In some
cases, dietary restrictions in humans can lead to deficiency diseases;
however, stable human groups have adapted to many dietary
patterns through both genetic specialization and cultural
conventions to use nutritionally balanced food sources.[220] The
human diet is prominently reflected in human culture and has led to
the development of food science.[221] Humans living in Bali, Indonesia,
preparing a meal
Until the development of agriculture, Homo sapiens employed a
hunter-gatherer method as their sole means of food collection.[221]
This involved combining stationary food sources (such as fruits, grains, tubers, and mushrooms, insect
larvae and aquatic mollusks) with wild game, which must be hunted and captured in order to be
consumed.[222] It has been proposed that humans have used fire to prepare and cook food since the time of
Homo erectus.[223] Human domestication of wild plants began about 11,700 years ago, leading to the
development of agriculture,[224] a gradual process called the Neolithic Revolution.[225] These dietary
changes may also have altered human biology; the spread of dairy farming provided a new and rich source
of food, leading to the evolution of the ability to digest lactose in some adults.[226][227] The types of food
consumed, and how they are prepared, have varied widely by time, location, and culture.[228][229]
In general, humans can survive for up to eight weeks without food, depending on stored body fat.[230]
Survival without water is usually limited to three or four days, with a maximum of one week.[231] In 2020 it
is estimated 9 million humans die every year from causes directly or indirectly related to starvation.[232][233]
Childhood malnutrition is also common and contributes to the global burden of disease.[234] However,
global food distribution is not even, and obesity among some human populations has increased rapidly,
leading to health complications and increased mortality in some developed and a few developing countries.
Worldwide, over one billion people are obese,[235] while in the United States 35% of people are obese,
leading to this being described as an "obesity epidemic."[236] Obesity is caused by consuming more calories
than are expended, so excessive weight gain is usually caused by an energy-dense diet.[235]
Biological variation
There is biological variation in the human species – with traits such
as blood type, genetic diseases, cranial features, facial features,
organ systems, eye color, hair color and texture, height and build,
and skin color varying across the globe. The typical height of an
adult human is between 1.4 and 1.9 m (4 ft 7 in and 6 ft 3 in),
although this varies significantly depending on sex, ethnic origin,
and family bloodlines.[237][238] Body size is partly determined by
genes and is also significantly influenced by environmental factors A Libyan, a Nubian, a Syrian, and an
such as diet, exercise, and sleep patterns.[239] Egyptian, drawing by an unknown
artist after a mural of the tomb of
There is evidence that populations have adapted genetically to Seti I
various external factors. The genes that allow adult humans to
digest lactose are present in high frequencies in populations that
have long histories of cattle domestication and are more dependent
on cow milk.[240] Sickle cell anemia, which may provide increased
resistance to malaria, is frequent in populations where malaria is
endemic.[241][242] Populations that have for a very long time
inhabited specific climates tend to have developed specific
phenotypes that are beneficial for those environments – short stature
and stocky build in cold regions, tall and lanky in hot regions, and
with high lung capacities or other adaptations at high altitudes.[243]
Some populations have evolved highly unique adaptations to very
specific environmental conditions, such as those advantageous to
ocean-dwelling lifestyles and freediving in the Bajau.[244]
Genetic research has demonstrated that human populations native to the African continent are the most
genetically diverse[263] and genetic diversity decreases with migratory distance from Africa, possibly the
result of bottlenecks during human migration.[264][265] These non-African populations acquired new
genetic inputs from local admixture with archaic populations and have much greater variation from
Neanderthals and Denisovans than is found in Africa,[183] though Neanderthal admixture into African
populations may be underestimated.[266] Furthermore, recent studies have found that populations in sub-
Saharan Africa, and particularly West Africa, have ancestral genetic variation which predates modern
humans and has been lost in most non-African populations. Some of this ancestry is thought to originate
from admixture with an unknown archaic hominin that diverged before the split of Neanderthals and
modern humans.[267][268]
Humans are a gonochoric species, meaning they are divided into male and female sexes.[269][270][271] The
greatest degree of genetic variation exists between males and females. While the nucleotide genetic
variation of individuals of the same sex across global populations is no greater than 0.1%–0.5%, the genetic
difference between males and females is between 1% and 2%. Males on average are 15% heavier and
15 cm (6 in) taller than females.[272][273] On average, men have about 40–50% more upper body strength
and 20–30% more lower body strength than women at the same weight, due to higher amounts of muscle
and larger muscle fibers.[274] Women generally have a higher body fat percentage than men.[275] Women
have lighter skin than men of the same population; this has been explained by a higher need for vitamin D
in females during pregnancy and lactation.[276] As there are chromosomal differences between females and
males, some X and Y chromosome-related conditions and disorders only affect either men or women.[277]
After allowing for body weight and volume, the male voice is usually an octave deeper than the female
voice.[278] Women have a longer life span in almost every population around the world.[279] There are
intersex conditions in the human population, however these are rare.[280]
Psychology
The human brain, the focal point of the central nervous system in humans, controls the peripheral nervous
system. In addition to controlling "lower", involuntary, or primarily autonomic activities such as respiration
and digestion, it is also the locus of "higher" order functioning such as thought, reasoning, and
abstraction.[281] These cognitive processes constitute the mind, and, along with their behavioral
consequences, are studied in the field of psychology.
Humans have a larger and more developed prefrontal cortex than other primates, the region of the brain
associated with higher cognition.[282] This has led humans to proclaim themselves to be more intelligent
than any other known species.[283] Objectively defining intelligence is difficult, with other animals adapting
senses and excelling in areas that humans are unable to.[284]
There are some traits that, although not strictly unique, do set
humans apart from other animals.[285] Humans may be the only
animals who have episodic memory and who can engage in
"mental time travel".[286] Even compared with other social animals,
humans have an unusually high degree of flexibility in their facial
expressions.[287] Humans are the only animals known to cry
emotional tears.[288] Humans are one of the few animals able to
self-recognize in mirror tests[289] and there is also debate over to
what extent humans are the only animals with a theory of
mind.[290]
During sleep humans dream, where they experience sensory images and sounds. Dreaming is stimulated by
the pons and mostly occurs during the REM phase of sleep.[292] The length of a dream can vary, from a
few seconds up to 30 minutes.[293] Humans have three to five dreams per night, and some may have up to
seven.[294] Dreamers are more likely to remember the dream if awakened during the REM phase. The
events in dreams are generally outside the control of the dreamer, with the exception of lucid dreaming,
where the dreamer is self-aware.[295] Dreams can at times make a creative thought occur or give a sense of
inspiration.[296]
The process of acquiring knowledge and understanding through thought, experience, and the senses is
known as cognition.[306] The human brain perceives the external world through the senses, and each
individual human is influenced greatly by his or her experiences, leading to subjective views of existence
and the passage of time.[307] The nature of thought is central to psychology and related fields. Cognitive
psychology studies cognition, the mental processes underlying behavior.[308] Largely focusing on the
development of the human mind through the life span, developmental psychology seeks to understand how
people come to perceive, understand, and act within the world and how these processes change as they
age.[309][310] This may focus on intellectual, cognitive, neural, social, or moral development. Psychologists
have developed intelligence tests and the concept of intelligence quotient in order to assess the relative
intelligence of human beings and study its distribution among population.[311]
Emotional experiences perceived as pleasant, such as joy, interest or contentment, contrast with those
perceived as unpleasant, like anxiety, sadness, anger, and despair.[322] Happiness, or the state of being
happy, is a human emotional condition. The definition of happiness is a common philosophical topic. Some
define it as experiencing the feeling of positive emotional affects, while avoiding the negative
ones.[323][324] Others see it as an appraisal of life satisfaction or quality of life.[325] Recent research
suggests that being happy might involve experiencing some negative emotions when humans feel they are
warranted.[326]
Love most commonly refers to a feeling of strong attraction or emotional attachment. It can be impersonal
(the love of an object, ideal, or strong political or spiritual connection) or interpersonal (love between
humans).[335] When in love dopamine, norepinephrine, serotonin and other chemicals stimulate the brain's
pleasure center, leading to side effects such as increased heart rate, loss of appetite and sleep, and an intense
feeling of excitement.[336]
Culture
Humanity's unprecedented set of intellectual skills were a key
Human society statistics
factor in the species' eventual technological advancement and
concomitant domination of the biosphere.[340] Disregarding extinct Most widely English,
hominids, humans are the only animals known to teach spoken Mandarin
[337][338]
generalizable information,[341] innately deploy recursive languages Chinese,
embedding to generate and communicate complex concepts,[342] Hindi,
Spanish,
engage in the "folk physics" required for competent tool
Standard
design,[343][344] or cook food in the wild.[345] Teaching and
Arabic,
learning preserves the cultural and ethnographic identity of human
Bengali,
societies.[346] Other traits and behaviors that are mostly unique to
French,
humans include starting fires,[347] phoneme structuring[348] and
Russian,
vocal learning.[349]
Portuguese,
Urdu
Language Most practiced Christianity,
While many species communicate, language is unique to humans, a religions[338][339] Islam,
defining feature of humanity, and a cultural universal.[350] Unlike Hinduism,
the limited systems of other animals, human language is open – an Buddhism, folk
infinite number of meanings can be produced by combining a religions,
limited number of symbols.[351][352] Human language also has the Sikhism,
capacity of displacement, using words to represent things and Judaism,
happenings that are not presently or locally occurring but reside in unaffiliated
the shared imagination of interlocutors.[149]
Language differs from other forms of
communication in that it is modality independent;
the same meanings can be conveyed through
different media, audibly in speech, visually by
sign language or writing, and through tactile
media such as braille.[353] Language is central to
the communication between humans, and to the
Principal language families of the world (and in some
sense of identity that unites nations, cultures and
cases geographic groups of families). For greater detail,
ethnic groups.[354] There are approximately six
see Distribution of languages in the world.
thousand different languages currently in use,
including sign languages, and many thousands
more that are extinct.[355]
The arts
Human arts can take many forms including visual, literary, and performing. Visual art can range from
paintings and sculptures to film, fashion design, and architecture.[356] Literary arts can include prose,
poetry, and dramas. The performing arts generally involve theatre, music, and dance.[357][358] Humans
often combine the different forms (for example, music videos).[359] Other entities that have been described
as having artistic qualities include food preparation, video games, and medicine.[360][361][362] As well as
providing entertainment and transferring knowledge, the arts are also used for political purposes.[363]
Evidence of humans engaging in musical activities predates cave art and so far music has been practiced by
virtually all known human cultures.[369] There exists a wide variety of music genres and ethnic musics;
with humans' musical abilities being related to other abilities, including complex social human
behaviours.[369] It has been shown that human brains respond to music by becoming synchronized with the
rhythm and beat, a process called entrainment.[370] Dance is also a form of human expression found in all
cultures[371] and may have evolved as a way to help early humans communicate.[372] Listening to music
and observing dance stimulates the orbitofrontal cortex and other pleasure sensing areas of the brain.[373]
Unlike speaking, reading and writing does not come naturally to humans and must be taught.[374] Still,
literature has been present before the invention of words and language, with 30,000-year-old paintings on
walls inside some caves portraying a series of dramatic scenes.[375] One of the oldest surviving works of
literature is the Epic of Gilgamesh, first engraved on ancient Babylonian tablets about 4,000 years ago.[376]
Beyond simply passing down knowledge, the use and sharing of imaginative fiction through stories might
have helped develop humans' capabilities for communication and increased the likelihood of securing a
mate.[377] Storytelling may also be used as a way to provide the audience with moral lessons and encourage
cooperation.[375]
China developed paper, the printing press, gunpowder, the compass and other important inventions.[385]
The continued improvements in smelting allowed forging of copper, bronze, iron and eventually steel,
which is used in railways, skyscrapers and many other products.[386] This coincided with the Industrial
Revolution, where the invention of automated machines brought major changes to humans' lifestyles.[387]
Modern technology is observed as progressing exponentially,[388] with major innovations in the 20th
century including: electricity, penicillin, semiconductors, internal combustion engines, the Internet, nitrogen
fixing fertilisers, airplanes, computers, automobiles, contraceptive pills, nuclear fission, the green revolution,
radio, scientific plant breeding, rockets, air conditioning, television and the assembly line.[389]
Philosophy is a field of study where humans seek to understand fundamental truths about themselves and
the world in which they live.[412] Philosophical inquiry has been a major feature in the development of
humans' intellectual history.[413] It has been described as the "no man's land" between definitive scientific
knowledge and dogmatic religious teachings.[414] Major fields of philosophy include metaphysics,
epistemology, logic, and axiology (which includes ethics and aesthetics).[415]
Society
Society is the system of organizations and institutions arising from interaction between humans. Humans are
highly social and tend to live in large complex social groups. They can be divided into different groups
according to their income, wealth, power, reputation and other factors. The structure of social stratification
and the degree of social mobility differs, especially between modern and
traditional societies.[416] Human groups range from the size of families to
nations. The first form of human social organization is thought to have
resembled hunter-gatherer band societies.[417]
Gender
Human societies typically exhibit gender identities and gender roles that
distinguish between masculine and feminine characteristics and prescribe
the range of acceptable behaviours and attitudes for their members based on
their sex.[418][419] The most common categorisation is a gender binary of
men and women.[420] Some societies recognise a third gender,[421] or less Humans often live in family-
based social structures
commonly a fourth or fifth.[422][423] In some other societies, non-binary is
used as an umbrella term for a range of gender identities that are not solely
male or female.[424]
Gender roles are often associated with a division of norms, practices, dress,
behavior, rights, duties, privileges, status, and power, with men enjoying
more rights and privileges than women in most societies, both today and in
the past.[425] As a social construct,[426] gender roles are not fixed and vary
historically within a society. Challenges to predominant gender norms have
recurred in many societies.[427][428] Little is known about gender roles in
the earliest human societies. Early modern humans probably had a range of
gender roles similar to that of modern cultures from at least the Upper
Paleolithic, while the Neanderthals were less sexually dimorphic and there
Depiction of a man and a
is evidence that the behavioural difference between males and females was
woman from the Pioneer
minimal.[429] plaque
Kinship
All human societies organize, recognize and classify types of social relationships based on relations
between parents, children and other descendants (consanguinity), and relations through marriage (affinity).
There is also a third type applied to godparents or adoptive children (fictive). These culturally defined
relationships are referred to as kinship. In many societies, it is one of the most important social organizing
principles and plays a role in transmitting status and inheritance.[430] All societies have rules of incest taboo,
according to which marriage between certain kinds of kin relations is prohibited, and some also have rules
of preferential marriage with certain kin relations.[431]
Ethnicity
Human ethnic groups are a social category that identifies together as a group based on shared attributes that
distinguish them from other groups. These can be a common set of traditions, ancestry, language, history,
society, culture, nation, religion, or social treatment within their residing area.[432][433] Ethnicity is separate
from the concept of race, which is based on physical characteristics, although both are socially
constructed.[434] Assigning ethnicity to a certain population is complicated, as even within common ethnic
designations there can be a diverse range of subgroups, and the makeup of these ethnic groups can change
over time at both the collective and individual level.[175] Also, there is no generally accepted definition of
what constitutes an ethnic group.[435] Ethnic groupings can play a powerful role in the social identity and
solidarity of ethnopolitical units. This has been closely tied to the rise of the nation state as the predominant
form of political organization in the 19th and 20th centuries.[436][437][438]
Governments create laws and policies that affect the citizens that they govern. There have been many forms
of government throughout human history, each having various means of obtaining power and the ability to
exert diverse controls on the population.[442] Approximately 47% of humans live in some form of a
democracy, 17% in a hybrid regime, and 37% in an authoritarian regime.[443] Many countries belong to
international organizations and alliances; the largest of these is the United Nations, with 193 member
states.[444]
Early human economies were more likely to be based around gift giving instead of a bartering system.[450]
Early money consisted of commodities; the oldest being in the form of cattle and the most widely used
being cowrie shells.[451] Money has since evolved into governmental issued coins, paper and electronic
money.[451] Human study of economics is a social science that looks at how societies distribute scarce
resources among different people.[452] There are massive inequalities in the division of wealth among
humans; the eight richest humans are worth the same monetary value as the poorest half of all the human
population.[453]
Conflict
Humans commit violence on other humans at a rate comparable to
other primates, but have an increased preference for killing adults,
infanticide being more common among other primates.[454]
Phylogenetic analysis predicts that 2% of early H. sapiens would be
murdered, rising to 12% during the medieval period, before
dropping to below 2% in modern times.[455] There is great variation
in violence between human populations, with rates of homicide
about 0.01% in societies that have legal systems and strong cultural
attitudes against violence.[456]
American troops landing at
The willingness of humans to kill other members of their species en Normandy, WWII.
masse through organized conflict (i.e., war) has long been the
subject of debate. One school of thought holds that war evolved as
a means to eliminate competitors, and has always been an innate human characteristic. Another suggests
that war is a relatively recent phenomenon and has appeared due to changing social conditions.[457] While
not settled, current evidence indicates warlike predispositions only became common about 10,000 years
ago, and in many places much more recently than that.[457] War has had a high cost on human life; it is
estimated that during the 20th century, between 167 million and 188 million people died as a result of
war.[458] War casualty data is less reliable for pre-medieval times, especially global figures. But compared
with any period over the past 600 years, the last ~80 years (post 1946), has seen a very significant drop in
global military and civilian death rates due to armed conflict. [459]
See also
Mammals portal
Evolutionary
biology portal
Science portal
Notes
1. The world population and population density statistics are updated automatically from a
template that uses the CIA World Factbook and United Nations World Population
Prospects.[118][119]
2. Cities with over 10 million inhabitants as of 2018.[120]
3. Traditionally this has been explained by conflicting evolutionary pressures involved in
bipedalism and encephalization (called the obstetrical dilemma), but recent research
suggest it might be more complicated than that.[197][198]
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