Vovides etal 2016

Explorations in central Veracruz on the Transvolcanic Mexican Belt and regions explored by 19th century botanists we found various populations of Ceratozamia whose taxonomic status is not yet completely clear. Especially two populations of C. mexicana with distinct morphology; one population at El Esquilón considered as C. mexicana by Chamberlain and other authors, and another population at El Mirador that Stevenson found a voucher at the herbarium in Paris with oblanceolate leaflets collected by Ghiesbrecht and designated it as Brongniart's holotype. The plants from El Mirador are comparable to the holotype of C. mexicana, but those at El Esquilón do not. Stevenson later also found a voucher at Kew by Thiselton-Dyer as C. mexicana var. tenuis from Mexico with linear-lanceolate leaflets that is similar to the plants at the El Esquilón population. An analysis of 11 leaflet anatomical characters gave a significant discriminant pattern (Wilk's lambda 0.00015 P < 0.01). The squared Mahalanobis distances were also significant (P < 0.05) and the first canonic variable showed that 93.4 % of the variation was due to the anatomical characters. A new nomenclatural combination is proposed and Thiselton-Dyer's voucher assigned as the lectotype of Ceratozamia tenuis. Epitypes were also designated for the two species in support of the anatomical analysis as well as a description for both species given.

Explorations in central Veracruz on the Transvolcanic Mexican Belt and regions explored by 19th century botanists we found various populations of Ceratozamia whose taxonomic status is not yet completely clear. Especially two populations of C. mexicana with distinct morphology; one population at El Esquilón considered as C. mexicana by Chamberlain and other authors, and another population at El Mirador that Stevenson found a voucher at the herbarium in Paris with oblanceolate leaflets collected by Ghiesbrecht and designated it as Brongniart's holotype. The plants from El Mirador are comparable to the holotype of C. mexicana, but those at El Esquilón do not. Stevenson later also found a voucher at Kew by Thisel-ton-Dyer as C. mexicana var. tenuis from Mexico with linear-lanceolate leaflets that is similar to the plants at the El Esquilón population. An analysis of 11 leaflet anatomical characters gave a significant discriminant pattern (Wilk's lambda 0.00015 P < 0.01). The squared Mahalanobis distances were also significant (P < 0.05) and the first canonic variable showed that 93.4 % of the variation was due to the anatomical characters. A new nomenclatural combination is proposed and Thiselton-Dyer's voucher assigned as the lectotype of Ceratozamia tenuis. Epitypes were also designated for the two species in support of the anatomical analysis as well as a description for both species given.

Botanical Sciences, 2016

Explorations in central Veracruz on the Transvolcanic Mexican Belt and regions explored by 19th century botanists we found various populations of Ceratozamia whose taxonomic status is not yet completely clear. Especially two populations of C. mexicana with distinct morphology; one population at El Esquilón considered as C. mexicana by Chamberlain and other authors, and another population at El Mirador that Stevenson found a voucher at the herbarium in Paris with oblanceolate leaflets collected by Ghiesbrecht and designated it as Brongniart’s holotype. The plants from El Mirador are comparable to the holotype of C. mexicana , but those at El Esquilón do not. Stevenson later also found a voucher at Kew by Thiselton-Dyer as C. mexicana var. tenuis from Mexico with linear-lanceolate leaflets that is similar to the plants at the El Esquilón population. An analysis of 11 leaflet anatomical characters gave a significant discriminant pattern (Wilk’s lambda 0.00015 P &lt; 0.01). The squared ...

Brittonia, 1982

Systematics and …, 2009

Brittonia, 1986

Mexico). A new species of Ceratozamia (Zamiaceae) from Veracruz, Mexico with comments on species relationships, habitats, and vegetative morphology in Ceratozamia. Brittonia 38: 17-26. 1986.-Ceratozamia euryphyllidia from Veracruz is described and illustrated. This species differs from all others in the genus in its very large, broad leaflets (9-16 cm wide) which have undulate margins, prominent veins, and a membranous to translucent texture at maturity. It is apparently related to a group of species which all have relatively small strobili and leaflets that are attenuate basally, oblanceolate to obovate, inequilateral, abruptly acuminate apically, and generally papyraceous to membranaceous. The allopatric species of this group inhabit moist to very wet areas. The wetter the habitat, the larger and thinner the leaflets. Thus, for example, C. latifolia which has the smallest leaflets grows in the driest habitat whereas C. euryphyllidia which grows in the wettest habitat has the largest leaflets in the genus. Ceratozamia euryphyllidia Vázquez Torres, Sabato & Stevenson (Figs. 1-5) Truncus semihypogaeus, humilis, ad 20 cm altus; cataphylla lanata, triangularia, 5 cm longa, basi 3.5 cm lata; folia pauca, usque 10, glabra; petiolus subteres teresve, 60-90 cm longus, parte infima dilatatus, validis spinis armatus; rachis semiteres, in dimidio inferiore paucis spinis armata, supra fere inermis vel inermis, in cuspidem 2-7 cm longam excurrens; foliola subopposita vel alterna, 6-13-juga, remota, oblanceolata vel late obovata, 18-31 cm longa, 9-16 cm lata, translucida, tenuia, basi cuneata, apicem versus subito asymmetrice caudato-acuminata, acumine fere 4 cm longo, margine integerrima, prope apicem repanda vel undulata, 40-66 nervis praedita; strobilus m lineari-cylindricus, 28 cm longus, 3 cm latus; pedunculus tomentosus, 6-8 cm longus, 1.5-2 cm latus; strobilus f cylindricus, 20 cm longus, 5 cm latus; pedunculus tomentosus, 12 cm longus, 1.4 cm latus; semina 2.3-2.7 cm longa, 2 cm lata.

Botanical Journal of the Linnean Society, 2004

Botanical Journal of the Linnean Society, 2008

Ceratozamia chimalapensis sp. nov. is described and illustrated. It is related to C. mirandae Vovides, Pérez-Farrera & Iglesias from Chiapas, but differs in trunk and peduncle size as well as in diameter of both megastrobili and microstrobili. Petiole, megasporophyll and indument colour also differ from that of C. mirandae. Ceratozamia chimalapensis forms part of the C. norstogii D.W.Stev. species complex, a group of ceratozamias with narrow leaflets growing in the herbaceous layer of oak forests in southern Mexico. These forests were severely affected by forest fires during 1998 and we recommend an IUCN Red List Category of CR B.1. Speciation in Ceratozamia has been discussed in the light of floristic refugia.

Brittonia, 1985

Premise of research. Many Ceratozamia species are closely related and form species complexes with great similarity among the component taxa, especially in juvenile stages. This, coupled with character loss in herbarium specimens, has resulted in debate regarding recognition and validation of the species. This study, focused on the four species comprising the Ceratozamia norstogii species complex from a morphological and leaflet anatomy viewpoint, was undertaken to clarify relationships among the four taxa in the complex.