Clifford Keil

Pontificia Universidad Católica del Ecuador, Museum of Invertebrates, Faculty Member

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Julian D Reynolds

Trinity College Dublin

James Elkins

School of the Art Institute of Chicago

David Brunsma

Virginia Tech

Kristen J Gremillion

Ohio State University

Eve Emshwiller

University of Wisconsin-Madison

Maurizio Forte

Duke University

David Seamon

Kansas State University

Jana Javornik

University of East London

Armando Marques-Guedes

UNL - New University of Lisbon

Gwen Robbins Schug

University of North Carolina at Greensboro

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Papers by Clifford Keil

C-band morphology of T(8;9)/8;9 in Blattella germanica

Theoretical and Applied Genetics, 1982

Centromere position and each arm of the T(8;9)/8;9 quadrivalent at late pachytene can be recogniz... more Centromere position and each arm of the T(8;9)/8;9 quadrivalent at late pachytene can be recognized by C-banding. The chromosome 9 breakpoint lies immediately adjacent to the centromeric C-band; that of 8, in the general region of the centromere but the relationship to centromeric bands was not determined since the latter stain very faintly. Chromosome 9 differs from no. 8 in the presence of a complex series of intercalary bands, heavy centromeric bands and, overall, a larger amount of C-band material. Possible implications of these differences with respect to chromosome breakability and the nature and distribution of mutant loci are noted. Earlier identification of adjacent-1 and adjacent-2 metaphase I ring orientations, made on the basis of orientation pattern, was confirmed.

Redescription of Pandora gloeospora (Zygomycetes: Entomophthorales) from Lycoriella mali (Diptera: Sciaridae)

Mycotaxon, 1990

An Analysis of Embryonic Trapping in the German Cockroach

Entomologia Experimentalis Et Applicata, Nov 1, 1977

The relationship between embryonic lethals from translocation heterozygotes and sterility from em... more The relationship between embryonic lethals from translocation heterozygotes and sterility from embryonic trapping in oothecae of the German cockroach has been analyzed. A modified constrained regression gives a means of predicting the expected hatch from lethality caused by high genetic loads, as in double translocation heterozygotes. Complete sterility is predicted at about 82.1% lethality.

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Identification and Characterization of a Permethrin Resistance Mechanism in Populations of the Fungus GnatLycoriella Mali(Fitch) (Diptera: Sciaridae)

Pesticide Biochemistry and Physiology, Jul 1, 1997

The commercial mushroom pest Lycoriella mali (Fitch) (Diptera: Sciaridae) has evolved resistance ... more The commercial mushroom pest Lycoriella mali (Fitch) (Diptera: Sciaridae) has evolved resistance to the adulticide permethrin. Preliminary investigations have indicated that oxidative metabolism was the most likely resistance mechanism. Monooxygenase activity (dealkylation reactions) against model substrates was used to examine specific metabolic activity associated with resistance. Differences in monooxygenase activity reflected resistance ranking of the field populations. The most resistant population was determined, by carbon monoxide difference spectrophotometry, to have a 1.7-fold greater level of cytochrome P-450 compared to the next most resistant population. This finding suggested that the quantity of enzyme was not solely responsible for variation in resistance. Enzyme kinetics were used to compare variation in metabolism of two model substrates, tetramethylbenzidine and para-phenylenediamine, between populations. The enzyme preparation from the most resistant population had a higher affinity for both model substrates. This suggested that this population possessed a different monooxygenase and that this enzyme was more active in metabolizing the model substrates. ᭧1997 Academic Press

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Insecticide Resistance

CRC Press eBooks, Mar 5, 2011

Genetic variability in the German cockroach

Journal of Heredity, Sep 1, 1978

Control of Adult Lycoriella mali and Megaselia halterata

Elsevier eBooks, 1987

Abstract Keil, C. B., 1986. Control of Adult Lycoriella mali and Megaselia halterata. It is impor... more Abstract Keil, C. B., 1986. Control of Adult Lycoriella mali and Megaselia halterata. It is important to adequately control adult Lycoriella mali and Megaselia halterata both to limit larval populations in compost and to reduce mechanical transmission of mushroom pathogens, particularly Verticillium fungicola. Evaluation of currently registered compounds and application techniques was undertaken in an effort to improve efficacy of control practices. Permethrin, dichlorvos, piperonyl butoxide and combinations were applied with a thermal fogger. Carrier was applied as a control. Permethrin was also evaluated in a dust formulation. Applications were made late in the crop cycle when populations were high. Evaluation of control was made with replicated light trap counts of adult flies 20 hours after application. Trap response to fly numbers was linear. Initial population size varied significantly and ranged from under ten flies of each species per minute to 3650 M . halterata and 78. 50 L. mali per minute. Two liquid formulations of permethrin, and permethrin dust gave variable results. Control of M. halterata never exceeded 80%. L. mali populations were reduced from 65% to almost 90%. The addition of piperonyl butoxide to the fog mix was not effective in increasing the toxicity of permethrin. The dust formulation and fog gave similar results at low populations but the fog was ineffective at higher fly levels. Dichlorvos was very effective as a fog and reduced population size by about 90% for both species. A permethrin-dichlorvos combination was very toxic to both species but not more so than dichlorvos alone. Sub-lethal dosages of dichlorvos disrupted attraction of both M. halterata and L. mali to chemical attractants. Permethrin resistance appears to be developing in mushroom fly populations, particularly in M. halterata. Large phorid populations are very difficult to control with this compound. Laboratory studies indicate that L. mali populations exhibit a range of tolerances to permethrin and may soon exhibit resistance in the field. Development of additional adulticides may be necessary to adequately control insects and Verticillium.

The Release of Sterile Males Into Natural Populations of the German Cockroach Blattella Germanica

Entomologia Experimentalis Et Applicata, Nov 1, 1981

A pilot experiment in genetic control of the German cockroach was conducted aboard a small vessel... more A pilot experiment in genetic control of the German cockroach was conducted aboard a small vessel based at Norfolk, Virginia. It utilized "sterile" ~ c~ (double translocation heterozygotes) whose sterility effects were due to embryonic lethality plus complete entrapment of any remaining viable embryos within the egg case. Initial infestations were reduced by insecticides prior to male release. Three releases of unmated ~ g were made at monthly intervals. Infested harborages, located in the course of insecticide application, were used as release sites. The experiment was terminated after 41/2 months. The results showed that released 6 ~ joined groups near release sites and that they competed well against wild type c~ g. Apparently neither they nor 9 9 with which they mated moved far from these sites. Sterility effects differed in respect to specific sites and general areas. Population growth was retarded markedly in the galley, the area of heaviest initial infestation. Terminal infestation was heaviest in the mess deck, although the highest sterility among 9 9 occurred at/near the mess-deck release site (only one mess deck-infested site was found prior to initation of the experiment). It is suggested that a slight increase in the number released would have suppressed/eliminated groups inhabiting galley harborages, but that site selection was the major problem in the mess deck. Analyses of nymphal age classes and mating types among 9 ? led to a hypothesis that insecticide-induced dispersal of nymphs resulted in the infestation of many new mess-deck harborages. Population growth was unchecked at these sites because they were too far removed from the sterile male release sites.

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Basic studies of chiasma frequency in male Blattella germanica (L.)

Apyrene Spermatogenesis in Ostrinia nubilalis (Hübner): Obligate and Facultative Diapausing Strains

Journal of Heredity, 1990

Four geographic strains of Ostrinia nubilalis (Hubner) differing in diapause characteristics were... more

Chromosome banding of German cockroach spermatocytes

Biological Control for Mushroom Production

Heinz, K.M. et. al. 2004. BioControl in Protected Culture. Ball Press, Batavia,, 2004

AN overview of mushroom production and an introduction to the primary insect and disease pests of... more AN overview of mushroom production and an introduction to the primary insect and disease pests of mushrooms is provided. The use of entomopathogenic nematodes, parasitoids, predatory insects andfungal pathogens is reviewed. Techniques for monitoring insect populations in mushroom houses and general pest management are explained.

Permethrin Resistance in Lycoriella mali (Diptera: Sciaridae)

Journal of Economic Entomology, 1989

ABSTRACT Four colonies of Lycoriella mali (Fitch) collected from different mushroom farms in sout... more ABSTRACT Four colonies of Lycoriella mali (Fitch) collected from different mushroom farms in southeastern Pennsylvania and Delaware were examined for resistance to topically applied permethrin in laboratory studies. Low to moderate (8- to 47-fold at LD50)resistance was discovered in all four colonies compared with a reference strain not previously exposed to permethrin. Resistance at the LD95 was from 34- to 67-fold. The LC95 of three of the colonies was approximately equal to the field rate for permethrin, whereas one colony&#39;s LC95 was below the field rate. Slopes of the dose-mortality lines of the four colonies ranged from 0.82 to 1.31, suggesting a potential for higher levels of resistance in these populations with continued selection pressure. The large concentration of mushroom farms in the area, short generation time of L. mali, numerous applications of permethrin, and history of pesticide use are likely to have influenced the development of resistance to permethrin in this species.

Cicadellinae of Ecuador and Cicadellidae of Galápagos

Neotropical Biodiversity, 2021

An annotated list of the Cicadellinae (Hemiptera: Auchenorrhyncha: Cicadellidae) of Ecuador is pr... more

Structure and Estimation of Shipboard German Cockroach ( Blattella germanica1 ) Populations

Environmental Entomology, 1981

ABSTRACT The removal method of population estimation was applied to populations of Blattella germ... more ABSTRACT The removal method of population estimation was applied to populations of Blattella germanica (Dictyoptera: Blattellidae) aboard two ships. A preponderance of adult females was collected from both ships. A low frequency of females with oothecae indicated that many were senescent or were delaying reproduction. Instar determination was made by head capsule measurements. Female nymphs appeared to have six instars, whereas males had only five. Structure of the two ships&#39; populations indicated that cockroaches were at near carrying capacity on the larger vessel, whereas the population on the smaller ship was less dense. Calculation of the removal method estimators for the two populations indicated that adult females are more resistant to capture than nymphs or adult males. Significant bias in estimation of population size resulted if this factor was not accounted for. Both precollection and residual population size was estimated with coefficients of variation of 3 to 13%.

FIGURES 74–76 in New species and host plants of Anastrepha (Diptera: Tephritidae) primarily from Suriname and Pará, Brazil

FIGURES 74–76. Eversible membranes, dorsal: 74, A. aithogaster (Suriname: Brownsberg, USNMENT0152... more

FIGURES 31–42 in New species and host plants of Anastrepha (Diptera: Tephritidae) primarily from Suriname and Pará, Brazil

FIGURES 31–42. Wings: 31–34, A. crassaculeus (Colombia: Pamplonita, ICAMF00000437; Suriname: Brow... more FIGURES 31–42. Wings: 31–34, A. crassaculeus (Colombia: Pamplonita, ICAMF00000437; Suriname: Brownsberg, USN- MENT00875104, USNMENT00875105, USNMENT00875106); 35–39, A. curvivenis (Peru: Tarapoto, USNMENT00744647; Suriname: Brownsberg, USNMENT00875205, Nassau, USNMENT00875206, USNMENT00875207, USNMENT00875208); 40–42, A. fuscoalata (Suriname: Berg en Dal, USNMENT00875198, Brownsberg, USNMENT00875200, USN- MENT00875197).

FIGURES 85–91 in New species and host plants of Anastrepha (Diptera: Tephritidae) primarily from Suriname and Pará, Brazil

FIGURES 85–91. Eversible membranes, dorsal unless otherwise indicated: 85–86, A. juxtalanceola (B... more FIGURES 85–91. Eversible membranes, dorsal unless otherwise indicated: 85–86, A. juxtalanceola (Brazil: REBIO-Rio Trombetas, USNMENT01113849; Suriname: Berg en Dal, USNMENT00875158), 87–88, A. microstrepha (Suriname: Brownsberg, USNMENT01354490; 87, dorsoapical); 89–90, A. neptis (Suriname: Brownsberg, USNMENT00875006; 90, dorsoapical); 91, A. sobrina (French Guiana: Mitaraka, USNMENT00875228).

FIGURES 17–18 in New species and host plants of Anastrepha (Diptera: Tephritidae) primarily from Suriname and Pará, Brazil

FIGURES 17–18. Habitus: 17, A. wachiperi (Peru: Villa Carmen, USNMENT01526566); 18, A. wittiensis... more

FIGURES 15–16 in New species and host plants of Anastrepha (Diptera: Tephritidae) primarily from Suriname and Pará, Brazil

FIGURES 15–16. Habitus: 15, A. tenebrosa (Peru: ExplorNapo, USNMENT00766120); 16, A. triangularis... more

C-band morphology of T(8;9)/8;9 in Blattella germanica

Theoretical and Applied Genetics, 1982

Redescription of Pandora gloeospora (Zygomycetes: Entomophthorales) from Lycoriella mali (Diptera: Sciaridae)

Mycotaxon, 1990

An Analysis of Embryonic Trapping in the German Cockroach

Entomologia Experimentalis Et Applicata, Nov 1, 1977

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Identification and Characterization of a Permethrin Resistance Mechanism in Populations of the Fungus GnatLycoriella Mali(Fitch) (Diptera: Sciaridae)

Pesticide Biochemistry and Physiology, Jul 1, 1997

Download

Insecticide Resistance

CRC Press eBooks, Mar 5, 2011

Genetic variability in the German cockroach

Journal of Heredity, Sep 1, 1978

Control of Adult Lycoriella mali and Megaselia halterata

Elsevier eBooks, 1987

The Release of Sterile Males Into Natural Populations of the German Cockroach Blattella Germanica

Entomologia Experimentalis Et Applicata, Nov 1, 1981

Download

Basic studies of chiasma frequency in male Blattella germanica (L.)

Apyrene Spermatogenesis in Ostrinia nubilalis (Hübner): Obligate and Facultative Diapausing Strains

Journal of Heredity, 1990

Four geographic strains of Ostrinia nubilalis (Hubner) differing in diapause characteristics were... more

Chromosome banding of German cockroach spermatocytes

Biological Control for Mushroom Production

Heinz, K.M. et. al. 2004. BioControl in Protected Culture. Ball Press, Batavia,, 2004

Permethrin Resistance in Lycoriella mali (Diptera: Sciaridae)

Journal of Economic Entomology, 1989

Cicadellinae of Ecuador and Cicadellidae of Galápagos

Neotropical Biodiversity, 2021

An annotated list of the Cicadellinae (Hemiptera: Auchenorrhyncha: Cicadellidae) of Ecuador is pr... more

Structure and Estimation of Shipboard German Cockroach ( Blattella germanica1 ) Populations

Environmental Entomology, 1981

FIGURES 74–76 in New species and host plants of Anastrepha (Diptera: Tephritidae) primarily from Suriname and Pará, Brazil

FIGURES 74–76. Eversible membranes, dorsal: 74, A. aithogaster (Suriname: Brownsberg, USNMENT0152... more

FIGURES 31–42 in New species and host plants of Anastrepha (Diptera: Tephritidae) primarily from Suriname and Pará, Brazil

FIGURES 85–91 in New species and host plants of Anastrepha (Diptera: Tephritidae) primarily from Suriname and Pará, Brazil

FIGURES 17–18 in New species and host plants of Anastrepha (Diptera: Tephritidae) primarily from Suriname and Pará, Brazil

FIGURES 17–18. Habitus: 17, A. wachiperi (Peru: Villa Carmen, USNMENT01526566); 18, A. wittiensis... more

FIGURES 15–16 in New species and host plants of Anastrepha (Diptera: Tephritidae) primarily from Suriname and Pará, Brazil

FIGURES 15–16. Habitus: 15, A. tenebrosa (Peru: ExplorNapo, USNMENT00766120); 16, A. triangularis... more

Clifford Keil

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