Papers by Walter Obermayer
More than 200 thalli of Sulcaria sulcata (lichenized Ascomycotina) from southeastern Tibet and ad... more More than 200 thalli of Sulcaria sulcata (lichenized Ascomycotina) from southeastern Tibet and adjacent regions have been analyzed chemically. Six major chemical races were detected, characterized by the occurrence of the following major compounds: psoromic acid (chemical race 1a), 2-methoxypsoromic acid (2a), 2-hydroxyvirensic acid (3a; often in combination with 2-hydroxyconvirensic acid), virensic acid (3b), and vulpinic acid, the latter in combination with either psoromic acid (1b) or 2-methoxypsoromic acid (2b).
Abstract: The holotype material of Lethariella sinensis Wei & Jiang is a mixture of two chemicall... more Abstract: The holotype material of Lethariella sinensis Wei & Jiang is a mixture of two chemically distinct taxa, one containing psoromic acid, the other containing norstictic acid. The former is chosen as lectotype and, as a result, L. mieheana Obermayer becomes a synonym of L. sinensis. Some further Lethariella specimens from Tibet have been subjected to TLC-analyses, and the results are presented.
Journal of The Hattori Botanical Laboratory, 1996

OBERMAYER, W. & KANTVILAS, G. 2003. The identity of the lichens Siphula himalayensis and Lecanora... more OBERMAYER, W. & KANTVILAS, G. 2003. The identity of the lichens Siphula himalayensis and Lecanora teretiuscula. - Herzogia 16: 27-34. The lichens Siphula himalayensis (Rasanen) Kantvilas and Lecanora teretiuscula Zahlbr., both recorded from the southeastern Himalayas and adjacent Tibetan regions, are regarded as conspecific with the widespread Arctic-alpine species Lecanora geophila (Th.Fr.) Poelt. The chemical constituents of this lichen include usnic acid, zeorin, traces of methylplacodiolic acid, some fatty acids and, frequently in Asian specimens, porphyrilic acid. The development of a fruticose habit by normally crustose genera and species growing under extreme environmental conditions is discussed briefly. Zusammenfassung: OBERMAYER, W. & KANTVILAS, G. 2003. Die Identitat der Flechten Siphula himalayensis und Lecanora teretiuscula. - Herzogia 16: 27-34. Die Flechtentaxa Siphula himalayensis (Rasanen) Kantvilas und Lecanora teretiuscula Zahlbr., die beide aus dem Sudost-Himalay...
Mycotaxon, 1994
ABSlRACT: Sb!: species of the genus Rinodina with thali andIOf discs PO+ orange due to the presen... more ABSlRACT: Sb!: species of the genus Rinodina with thali andIOf discs PO+ orange due to the presence of pannarin are hlf1eIto recognized in Eurasia. Ooe is saxicolous, R. sarfloOO9nsis, and live are corticolous, R. datnatica, R. eftIot'escens, R. &Xt:tBSCenS, R. granuIans and R. pnMeIJa. Notes on lhe IIDCOI'lomy 01 R. granulans are pf"O'llkKld. Descriptions and illustrations of this species and lhe also poor1y oodemood R. BXCr6SC6f1S are included. A key 10 the treated species is given. Other known specles contalning pannarin are briefIy discussed.

15 specimens of Buellia centralis and 106 specimens of Dimelaena oreina from Tibet and other Cent... more 15 specimens of Buellia centralis and 106 specimens of Dimelaena oreina from Tibet and other Central Asian regions have been studied chemically. Buellia centralis contains rhizocarpic, hypoprotocetraric and nornotatic acid. Molecular analyses revealed its taxonomic position within the core group of Buellia. Except for chemotype IV (gyrophoric acid and fumarprotocetraric acid) and VI (sphaerophorin and gyrophoric acid), all hitherto known chemotypes of Dimelaena oreina are present in the study area. Due to the occurrence of hypostictic acid (accompanied by stictic acid) as a major compound in Dimelaena oreina, a new chemotype (Vb) is defined. The use of just a single areola for TLC-investigations confirmed this new chemotype as well as the existence of chemotypes IV (gyrophoric acid and fumarprotocetraric acid), chemotype VII (stictic acid and gyrophoric acid), and the newly recognised (sub-)chemotypes of chemotype II (chemotype IIa with gyrophoric acid and chemotype IIb with ovoic a...

Zusammenfassung: Die Arten der Gattung Arthrorhaphis Th. Fr. werden für Europa und Gränland revid... more Zusammenfassung: Die Arten der Gattung Arthrorhaphis Th. Fr. werden für Europa und Gränland revidiert. Es werden fünf auf Artniveau stehende Sippen anerkannt: Arthrorhaphis a/pina (Schaerer) R. Santesson in D. Hawksworth, P . James & Coppins, A . citrinella (Acharius) Poelt, A . grisea Th. Fries, A . muddii W. Obermayer und A. vacillans Th. Fries & Almquist ex Th. Fries. Arthrorhaphis juscoreagens (Vainio) Poelt wird mit A. grisea synonymisiert, Arthrorhaphis muddii W. Obermayer (bisher häufig als A. juscoreagens bestimmt) wird als eine auf Dibaeis baeomyces (Syn.: Baeomyces roseus} wachsende Sippe neu beschrieben . Von Arthrorhaphis a/pina, A . citrinella und A . grisea werden Lectotypen ausgewählt. Das Auftreten von Calcium-Oxalat-Kristallen in der Medulla erweist sich als ein wichtiges diagnostisches Differentialmerkmal, das auch sterile Lager von Arthrorhaphis alpina und A. cacillans (mit Kristallen) gegen A. citrinella-Thalli (ohne Medullarkristalle) abgrenzen läßt. -Der Befall...
Summary: The distribution of the genus Anaptychia, Heterodermia, Hyperphyscia and Phaeophyscia (P... more Summary: The distribution of the genus Anaptychia, Heterodermia, Hyperphyscia and Phaeophyscia (Physciaceae) in Austria is presented in form of grid maps. The patterns of distribution are explicable by the description of the ecological range of the treated species. Kokkuvote: Perekondade Anaptychia, Heterodermia, Hyperphyscia jaPhaeophyscia (Physciaceae) levik Austrias. Perekondade Anaptychia, Heterodermia, Hyperphyscia jaPhaeophyscia (Physciaceae) levik Austrias on esitatud kaartidel. Vaadeldud liikide levikumustreid selgitatakse nende liikide okoloogiliste noudluste kirjeldamise abil.

Hafellner, J. & Obermayer, W. 2009. The role of Paranectria oropensis in community dynamics of ep... more Hafellner, J. & Obermayer, W. 2009. The role of Paranectria oropensis in community dynamics of epiphyte synusia on roadside trees. - Herzogia 22: 177-190. It is documented from a study site in the city of Graz (Austria, Styria) that beside the corticioid basidiomycete Athelia arachnoidea, the lichenicolous hypocrealean pyrenomycete Paranectria oropensis can play a comparably important role in opening lichen synusia on bark. Shortly after the death and the splitting off of the remnants of lichen thalli caused by an invasion with Paranectria oropensis, a recolonisation of the bare bark areas can be observed. Zusammenfassung: Hafellner, J. & Obermayer, W. 2009. Die Rolle von Paranectria oropensis in der Gesellschaftsdynamik von Epiphytensynusien an Alleebaumen. - Herzogia 22: 177-190. In einer in Graz (Osterreich, Steiermark) durchgefuhrten Studie wird gezeigt, dass der hypocreale Pyrenomyzet Paranectria oropensis bei der Offnung von borkenbewohnenden Flechtensynusien eine ahnlich bede...

Umbilicaria orientalis Davydov sp. nov. is described and phylogenetic analysis (ITS, mtLSU and RP... more Umbilicaria orientalis Davydov sp. nov. is described and phylogenetic analysis (ITS, mtLSU and RPB2) confirmed its distinctness and indicated a sister relationship with U. trabeculata within the U. vellea group. The species is morphologically similar to U. vellea but differs by simple, cylindrical or strap-like rhizinomorphs and by thalloconidia, developing both on the lower surface and on rhizinomorphs, that are 1–2 (rarely 4)–celled or in aggregates of up to 5–6 cells. Umbilicaria orientalis is described from the Russian Far East and is currently known from a wide range of localities in East Asia, from the Far East to South Siberia and Mongolia in the north, to Hebei and Tibet in the south. New sequences of U. americana were obtained; the species represents an independent phylogenetic lineage within Umbilicaria subg. Papillophora. Diagnostic traits and variability of different developmental stages of U. orientalis, as well as its East Asian distribution pattern, are discussed.
A floristic survey of the yellow cetrarioid lichens containing usnic acid (fam. Parmeliaceae, lic... more A floristic survey of the yellow cetrarioid lichens containing usnic acid (fam. Parmeliaceae, lichenized Ascomycota) which are found in Tibet and neighbouring areas is presented. Typical characters, chemical constituents and distribution data are reported for all 26 species from seven genera (Allocetraria, Cetraria, Cetreliopsis, Flavocetraria, Nephromopsis, Tuckneraria and Vulpicida). Complete descriptions are provided for several taxa which have not been included in recent reviews of the cetrarioid genera. Two identification keys for taxa - both at generic and species level - have been compiled. Worldwide distribution maps are presented for the first time for some Allocetraria, Cetraria, Cetreliopsis and Tuckneraria species.

Herzogia
Obermayer W., Witzmann M. & McCune B. 2018: Hypogymnia caperatica, a new species from the Hengdua... more Obermayer W., Witzmann M. & McCune B. 2018: Hypogymnia caperatica, a new species from the Hengduan Shan area (easternmost Tibetan region), with notes on some rare taxa from the greater Tibetan region and Bhutan.-Herzogia 31: 677-694. Hypogymnia caperatica, which contains caperatic acid as a major lichen substance in the medulla, is described as new from the Hengduan Shan area (southeasternmost boundaries of the Tibetan plateau). The aliphatic acid caperatic acid is reported for the first time from the genus Hypogymnia. The occurrence of fatty acids in the genus Hypogymnia is generally discussed. The very rare lichen Hypogymnia sikkimensis, hitherto only known from two localities in Sikkim, as well as Hypogymnia diffractaica and H. hengduanensis are reported as new for the Kingdom of Bhutan. Hypogymnia capitata, known from the type and paratype specimens only, is reported from three further localities in the East Tibetan region.

15 specimens of Buellia centralis and 106 specimens of Dimelaena oreina from Tibet and other Cent... more 15 specimens of Buellia centralis and 106 specimens of Dimelaena oreina from Tibet and other Central Asian regions have been studied chemically. Buellia centralis contains rhizo-carpic, hypoprotocetraric and nornotatic acid. Molecular analyses revealed its taxonomic posi-tion within the core group of Buellia. Except for chemotype IV (gyrophoric acid and fumar-protocetraric acid) and VI (sphaerophorin and gyrophoric acid), all hitherto known chemo-types of Dimelaena oreina are present in the study area. Due to the occurrence of hypostictic acid (accompanied by stictic acid) as a major compound in Dimelaena oreina, a new chemo-type (Vb) is defined. The use of just a single areola for TLC-investigations confirmed this new chemotype as well as the existence of chemotypes IV (gyrophoric acid and fumarprotocetraric acid), chemotype VII (stictic acid and gyrophoric acid), and the newly recognised (sub-)chem-otypes of chemotype II (chemotype IIa with gyrophoric acid and chemotype IIb with o...
Australian Journal of Chemistry
The morphology and the chemical constituents of the lichen Ramalina dumeticola from Fraser's Hill... more The morphology and the chemical constituents of the lichen Ramalina dumeticola from Fraser's Hill are described. This lichen is a new record for Malaysia.

MycoKeys
Based on the analysis of both historical and recent collections, this paper reports an annotated ... more Based on the analysis of both historical and recent collections, this paper reports an annotated list of taxa which are new to the lichen biota of Italy or of its administrative regions. Specimens were identified using a dissecting and a compound microscope; routine chemical spot tests and standardized thin-layer chromatography (TLC or HPTLC). The list includes 225 records of 153 taxa. Twenty taxa are new to Italy, the others are new to one or more administrative regions, with 15 second records and 5 third records for Italy. Some of the species belong to recently-described taxa, others are poorly known, sterile or ephemeral lichens which were largely overlooked in Italy. Several species are actually rare, either because of the rarity of their habitats (e.g. old-growth forests), or because in Italy they are at the margins of their bioclimatic distribution. The picture of the lichen biota of Italy has now new pixels, but its grain is still coarse. Further analysis of historical collec...

Plant and Fungal Systematics
The ‘competition-relatedness’ hypothesis postulates that co-occurring taxa should be more distant... more The ‘competition-relatedness’ hypothesis postulates that co-occurring taxa should be more distantly related, because of lower competition. This hypothesis has been criticized for its dependence on untested assumptions and its exclusion of other assembly forces beyond competition and habitat filtering to explain the co-existence of closely related taxa. Here we analyzed the patterns of co-occurring individuals of lichenized fungi in the Graphis scripta complex, a monophyletic group of species occurring in temperate forests throughout the Northern Hemisphere. We generated sequences for three nuclear ribosomal and protein markers (nuLSU, RPB2, EF-1) and combined them with previously generated sequences to reconstruct an updated phylogeny for the complex. The resulting phylogeny was used to determine the patterns of co-occurrences at regional and at sample (tree) scales by calculating standard effect size of mean pairwise distance (SES.MPD) among co-occurring samples to determine whethe...
Uploads
Papers by Walter Obermayer