
Karma Orong
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Papers by Karma Orong
simulate a precipitation decline, throughfall-exclusion (TFE) shelters were applied during three consecutive
monsoon seasons in an oak forest (2.650 m a.s.l.) and a conifer-dominated forest (3.260 m a.s.l.) in central
Bhutan. Leaf water potentials, tree mortality, stem increment, soil CO
2
efflux, litterfall and fine root dynamics
were assessed. TFE significantly and consistently decreased topsoil (0–30 cm) moisture and leaf water potentials
of Quercus lanata and Quercus griffithii (lower elevation), and to a lesser extend those of Tsuga dumosa and Quercus
semecarpifolia, (higher elevation). TFE did not impose tree mortality. Stem increment remained unaffected until
the second TFE year, but showed reductions during the third year with Tsuga dumosa being most severely affected
(-60%). Standing fine root biomass stocks were hardly affected by TFE. Increased root necromass and faster fine
root growth in the lower elevation forest suggest that the oak trees increased C allocation below ground. Soil CO
2
efflux sharply declined during all three TFE years in both forests. Above ground litter input was unaffected by
TFE until the second treatment year. Overall, both forest ecosystems appeared highly resistant to the imposed soil
drying, with no signs of tree mortality and stable living root biomass stocks.
simulate a precipitation decline, throughfall-exclusion (TFE) shelters were applied during three consecutive
monsoon seasons in an oak forest (2.650 m a.s.l.) and a conifer-dominated forest (3.260 m a.s.l.) in central
Bhutan. Leaf water potentials, tree mortality, stem increment, soil CO
2
efflux, litterfall and fine root dynamics
were assessed. TFE significantly and consistently decreased topsoil (0–30 cm) moisture and leaf water potentials
of Quercus lanata and Quercus griffithii (lower elevation), and to a lesser extend those of Tsuga dumosa and Quercus
semecarpifolia, (higher elevation). TFE did not impose tree mortality. Stem increment remained unaffected until
the second TFE year, but showed reductions during the third year with Tsuga dumosa being most severely affected
(-60%). Standing fine root biomass stocks were hardly affected by TFE. Increased root necromass and faster fine
root growth in the lower elevation forest suggest that the oak trees increased C allocation below ground. Soil CO
2
efflux sharply declined during all three TFE years in both forests. Above ground litter input was unaffected by
TFE until the second treatment year. Overall, both forest ecosystems appeared highly resistant to the imposed soil
drying, with no signs of tree mortality and stable living root biomass stocks.