Papers by José Rances Caicedo Portilla

<i>Atractus alphonsehogei</i> Cunha &amp; Nascimento, 1983 Fig. 8 <i>Atract... more <i>Atractus alphonsehogei</i> Cunha &amp; Nascimento, 1983 Fig. 8 <i>Atractus alphonsehogei</i> CUNHA &amp; NASCIMENTO, 1983:25 (FIG. 2 PP. 27). <b>Holotype.</b> Adult male (MPEG 14928) collected by O. Cunha and F. Nascimento on August 0 1 1978 at Bela Vista, Km 75 of the PA-242 road near municipality Nova Timboetuba (ca., 0 1 <i>°</i> 12'47''S, 47 <i>°</i> 23'18''W; 30 m asl), state of Pará, Brazil (Fig. 8). <b>Paratypes.</b> Thirteen specimens, eleven of them from the state of Pará and the other two from the state of Maranhão, all of them collected by O. Cunha, F. Nascimento or by both in the same fieldtrip: female (MPEG 2221) collected on September 20 1972 on road PA-140 from Santo Antônio do Tauá (ca., 0 1 <i>°</i> 08'S, 48 <i>°</i> 07'W; 20m asl) toward municipality of Vigia (00 <i>°</i> 51'S, 48 <i>°</i> 08'W; 10m asl); male (MPEG 8573) collected on March 0 2 1975 and female (MPEG 8667) collected on October 0 7 1974 on Parada Bom Jesus on the PA-242 road, 11 Km to municipality of Bragança (01 <i>°</i> 03'13''S, 46 <i>°</i> 45'56''W; 10m asl); male (MPEG 12593) collected on September 17 1976 at PA-140 road in the locality of Santa Rosa da Vigia 12 Km (airline) from the municipality of Vigia; female (MPEG 9949) collected on June 26 1975 at Cacoal Farm, Arari road a confluence of PA-242 27 Km from the municipality of Bragança; two males (MPEG 2976) and (MPEG 10130) collected on March 14 and 10 1976, respectively and three females (MPEG 6408) collected on March 27 1974, and (MPEG 10129) and (MPEG 10132) both collected on March 10 1976 at locality of Colônia Nova, Km 264 of the BR-316 highway near to Rio Gurupi (ca., 0 1 <i>°</i> 49'S, 48 <i>°</i> 24'W; 30m asl); female (MPEG 10093) collected on June 23 1975 at Km 224 of the BR-316 highway; and two other males from the state of Maranhão: (MPEG 10874) collected on November 0 2 1975 on BR- 316 highway 25 Km before Rio Gurupi (ca., 0 1 <i>°</i> 49'S, 46 <i>°</i> 06'W; 46m asl), Nova Vida, and (MPEG 11145) collected on February 23 1976 at São Raimundo ca. 8 Km from [...]

<i>Atractus</i> <i>collaris</i> Peracca, 1897 Figures 2, 3A–C, 4–6 <i&... more <i>Atractus</i> <i>collaris</i> Peracca, 1897 Figures 2, 3A–C, 4–6 <i>Atractus</i> <i>collaris</i> PERACCA 1897: 4 (CONONACO RIVER, PASTAZA, ECUADOR). <i>Leptocalamus limitaneus</i> AMARAL 1935: 219 (LA PEDRERA, AMAZONAS, COLOMBIA; IBSP 9196). <b>New synonymy.</b> [<i>Atractus</i>] <i>limitaneus</i> —SAVAGE 1960: 81. <i>Atractus collaris collaris—</i> DIxON &amp; SOINI 1977: 34 (IN PART). <i>Atractus alphonsehogei—</i> MARTINS &amp; OLIVEIRA 1993: 24 (FMT 450, A MALE SPECIMEN). <i>Atractus collaris</i> —CARRILLO &amp; ICOCHEA 1995: 13. <b>Holotype.</b> Adult male housed in the Museo Regionali di Scienza Naturali di Torino (MRSNT) from Cononaco River (c.a., 0 1 <i>°</i> 31'S, 75 <i>°</i> 36'W; 190m asl), province of Pastaza, Ecuador. Specimen apparently lost (see Andreaone &amp; Gavetti 2007) (Fig. 4). <b>Diagnosis.</b> <i>Atractus collaris</i> is distinguished from all congeners, except for those species of the <i>A. collaris</i> species group by having one (usually) or two (rarely) apical pits on dorsal scales from both sexes and supracloacal tubercles in the cloacal region of mature males (Passos <i>et al</i>. 2013b). Additionally, the following combination of morphological characters is unique of the species and distinguishes it from any other species of <i>Atractus</i>: (1) dorsal scale rows 17/17/17 with apical pits in both sexes and supracloacal tubercles in males; (2) postoculars two; (3) moderately long loreal, contacting first three supralabials; (4) temporals 1+2; (5) seven supralabials, third and fourth contacting eye; (6) seven infralabials, first three contacting chinshields; (7) five (rarely) or six maxillary teeth; (8) gular scale rows in four series; (9) preventrals usually four; (10) ventrals 167–186 in females, 145–178 in males; (11) subcaudals 18–24 in females, 22–33 in males; (12) in preservative, dorsum brown to grayish black, with cream occipital collar incomplete and small paired black spots usually cream bordered along the body, first dorsal scale rows with lighter center and brown lateral [...]
FIGURE 13. DISTRIBUTION OF Atractus alphonsehogei.THE BLACK CIRCLE REPRESENTS THE TYPE-LOCALITY.
FIGURE 12. DISTRIBUTION OF Atractus collaris (SqUARES) A. gaigeae (TRIANGLES). WE USE RECORDS BAS... more FIGURE 12. DISTRIBUTION OF Atractus collaris (SqUARES) A. gaigeae (TRIANGLES). WE USE RECORDS BASED ON THE SPECIMENS PRESERVED AT SCIENTIFIC COLLECTION AS WELL AS THE PHOTOGRAPHIC RECORDS WITH DETAILED PROVENANCE DATA (WHITE SqUARES WITH CENTRAL DOTS).THE BLACK SYMBOL SqUARE REPRESENTS THE TYPE-LOCALITY OF A.collaris.
FIGURE 11. BIVARIATE PLOTS WITH 95% CONFIDENCE REGIONS FOR THE FIRST TWO AxES DERIVED FROM SCORES... more FIGURE 11. BIVARIATE PLOTS WITH 95% CONFIDENCE REGIONS FOR THE FIRST TWO AxES DERIVED FROM SCORES OF DISCRIMINANT ANALYSES (DFA) WITH 1,000 REPLICATES OF BOOTSTRAP FOR MALES (A) AND FEMALES (B) FROM THE Atractus collaris COMPLEx.THE DIRECT COSINES DISPLAY THE VARIABLES MORE RELATED WITH THE DISCRIMINANT FUNCTIONS.
FIGURE 10. HEMIPENIAL MORPHOLOGY (DETAILED LINE ARTS ON THE LEFT AND PHOTOGRAPHS ON THE RIGHT) OF... more FIGURE 10. HEMIPENIAL MORPHOLOGY (DETAILED LINE ARTS ON THE LEFT AND PHOTOGRAPHS ON THE RIGHT) OF Atractus collaris (A, MNRJ 24959) FROM LAGUNA GRANDE SCIENTIFIC STATION, MUNICIPALITY OF CUYABENO, SUCUMBíOS, ECUADOR; Atractus gaigeae (B, USNM 217627) FROM MUNICIPALITY OF MONTALVO,PROVINCE OF PASTAZA,ECUADOR;AND Atractus alphonsehogei (C, MPEG 10874) FROM SãO RAIMUNDO,8 KM FROM MUNICIPALITY OF SANTA INÊS, STATE OF MARANHãO,BRAZIL.SCALE BAR = 1 MM.
FIGURE 9. MICRODERMATOGLYPHICS OF THE DORSAL SCALES OF Atractus collaris (A–B, MNRJ 24959) FROM L... more FIGURE 9. MICRODERMATOGLYPHICS OF THE DORSAL SCALES OF Atractus collaris (A–B, MNRJ 24959) FROM LAGUNA GRANDE SCIENTIFIC STATION,MUNICIPALITY OF CUYABENO,SUCUMBíOS,ECUADOR;Atractus gaigeae (C–D, USNM 217623)FROM MUNICIPALITY OF MONTALVO,PROVINCE OF PASTAZA,ECUADOR;AND Atractus alphonsehogei (E–F, MPEG 10874) FROM SãO RAIMUNDO,8 KM FROM MUNICIPALITY OF SANTA INÊS, STATE OF MARANHãO, BRAZIL. EACH SCALE REGION WAS PHOTOGRAPHED FROM BASAL PORTION (LEFT SIDE) AND APICAL PORTION (RIGHT SIDE).
FIGURE 7. DORSAL (A), LATERAL (B) AND VENTRAL (C) VIEWS OF HEAD,AND DORSAL (D) AND LATERAL (E)VIE... more FIGURE 7. DORSAL (A), LATERAL (B) AND VENTRAL (C) VIEWS OF HEAD,AND DORSAL (D) AND LATERAL (E)VIEWS OF BODY OF THE HOLOTYPE OF Atractus gaigeae (UMMZ 82887) FROM THE PROVINCE OF ZAMORA-CHINCHIPE,ECUADOR WITHOUT PRECISE LOCATION.
FIGURE 6. DORSAL (A), LATERAL (B) AND VENTRAL (C) VIEWS OF BODY OF THE HOLOTYPE OF Leptocalamus l... more FIGURE 6. DORSAL (A), LATERAL (B) AND VENTRAL (C) VIEWS OF BODY OF THE HOLOTYPE OF Leptocalamus limitaneus (IBSP 9196) FROM LA PEDRERA,MUNICIPALITY OF PUERTO CóRDOBA, DEPARTMENT OF AMAZONAS, COLOMBIA.PICTURE MODIFIED FROM AMARAL (1935). ARROW SHOWS A DISTINCT POSTNASAL SCALE (SEE TExT FOR DETAILS).
FIGURE4. DORSALVIEW OF THE ANTERIORPORTION OF BODY OF THE HOLOTYPEOF Atractus collaris FROM THE C... more FIGURE4. DORSALVIEW OF THE ANTERIORPORTION OF BODY OF THE HOLOTYPEOF Atractus collaris FROM THE CONONACORIVER,ECUADOR. PICTURE MODIFIEDFROM PERACCA (1897).
FIGURE 3. GENERAL VIEW IN LIFE OF Atractus collaris (A–B) BOTH SPECIMENS FROM MADRE SELVA, OROSA ... more FIGURE 3. GENERAL VIEW IN LIFE OF Atractus collaris (A–B) BOTH SPECIMENS FROM MADRE SELVA, OROSA RIVER AND A THIRD INDIVIDUAL (C) FROM SANTA CRUZ, MAZAN RIVER, BOTH LOCALITIES IN THE IqUITOS REGION, DEPARTMENT OF LORETO, PERU; AND Atractus gaigeae (D, QCAZ 8339) FROM COMUNIDAD TARANGARO,CAMPO VILLANO, PROVINCE OF PASTAZA,ECUADOR.PHOTOS A–C BY MATTEW CAGE (SPECIMENS NOT PRESERVED IN SCIENTIFIC COLLECTIONS) AND PHOTO D BY DIEGO PAUCAR AND REPTILIAWEBECUADOR.
FIGURE 2. UNCOLLECTED SPECIMEN OF Atractus collaris FROM NAUTA-IqUITOS ROAD,IN LORETO,PERU,ExHIBI... more FIGURE 2. UNCOLLECTED SPECIMEN OF Atractus collaris FROM NAUTA-IqUITOS ROAD,IN LORETO,PERU,ExHIBITING A DISTINCT VERTEBRAL STRIPE (ABSENT IN OTHER SPECIMENS FROM THE REGION),DEMONSTRATING LOCAL POLYMORPHISM OF THIS FEATURE IN THE SPECIES. PHOTOS BY KONRAD MEBERT.
FIGURE 1. DISTRIBUTION OF Atractus alphonsehogei (CIRCLES),A. collaris (SqUARES) AND A. gaigeae (... more FIGURE 1. DISTRIBUTION OF Atractus alphonsehogei (CIRCLES),A. collaris (SqUARES) AND A. gaigeae (TRIANGLES).
FIGURE 2. Dorsal (A), lateral (B), and ventral (C) views of head of the paratype of Tricheilostom... more FIGURE 2. Dorsal (A), lateral (B), and ventral (C) views of head of the paratype of Tricheilostoma brevissimum (MCZ 38950).
FIGURE 8. DORSAL (A), LATERAL (B) AND VENTRAL (C) VIEWS OF HEAD OF THE HOLOTYPE OF Atractus alpho... more FIGURE 8. DORSAL (A), LATERAL (B) AND VENTRAL (C) VIEWS OF HEAD OF THE HOLOTYPE OF Atractus alphonsehogei (MPEG 14928) FROM BELA VISTA,KM 75 PA-242 ROAD,STATE OF PARa,BRAZIL.PICTURE MODIFIED FROM CUNHA AND NASCIMENTO (1983).SCALE BAR = 4 MM.

Revista de la Academia Colombiana de Ciencias Exactas, Físicas y Naturales, 2021
The study of the introduction of exotic species in Colombia has been a trending topic in the last... more The study of the introduction of exotic species in Colombia has been a trending topic in the last decade due to their effects on local diversity. The records of exotic reptile species of the genus Hemidactylus have increased based on museum vouchers and fieldwork. However, there are still plenty of information gaps regarding their current distribution in the country. Similarly, there is limited information on unusual reptile distributions caused by human transport inside the country. Here, we present an update to the distribution of the exotic house geckos Hemidactylus frenatus, H. garnotii, and H. mabouia based on several new localities and historical records. We also present an atypical record of the golden spectacled tegu Gymnophthalmus speciosus in paramo ecosystems from the Central Andes of Colombia, almost 2000 meters above the upper elevational known limit for this species. The distribution update shows that H. frenatus is widely distributed in 30 of the 32 departments of Col...

Journal of Herpetology, 2012
are two particularly rare and poorly known lizards described from single specimens. In the case o... more are two particularly rare and poorly known lizards described from single specimens. In the case of A. pariaensis, it remains known from the holotype, whereas A. steyeri is known from three additional specimens reported in the literature after the original description of the species. A single new specimen of A. pariaensis and five of A. steyeri, including the first adult males recorded for both species, make possible a more representative description of both species, including descriptions of the hemipenes. Despite both species presenting some similar morphological characteristics, the examination of the hemipenial morphology revealed very different organs. The hemipenis of A. steyeri presents some characteristics that resemble the organs of two species from the Santa Marta Mountain Range in the ''bitaeniata-group'' (Anadia pulchella and Anadia altaserrania). On the other hand, the hemipenes of A. pariaensis are unique morphologically and cannot be associated with the hemipenes known from other species in the genus. We describe variation within both species, and we comment on possible sexual dimorphism (number and arrangement of the femoral pores), natural history, and the known geographic distribution of the species. We also comment on Anadia bumanguesa Rueda-Almonacid and Caicedo 2004 based on a new specimen, the second known. This species may be a synonym of A. steyeri.

Zootaxa, 2021
The genus Thamnodynastes is the most diverse within the tribe Tachymenini, with an extensive and ... more The genus Thamnodynastes is the most diverse within the tribe Tachymenini, with an extensive and complex taxonomic history. The brief descriptions and lack of robust diagnostic characters are the main sources for identification errors and for the difficulty to assess the diversity estimates of the genus. The Thamnodynastes pallidus group was briefly designated to encompass the most arboreal species of the genus, with thinner bodies and longer tails: T. pallidus, T. longicaudus, T. sertanejo, and a fourth undescribed species. After its designation, no other paper addressed this group and its morphological variation, especially for the hemipenis, is still undetermined. After the analysis of all species of Thamnodynastes we were able to corroborate the distinctiveness of the T. pallidus group and to accurately diagnose its fourth species from the western portion of the Amazonia lowlands. The new species is distinguishable from all congeners, except T. sertanejo, by the absence of ventr...

Caldasia
RESUMEN Se presenta nueva información sobre la taxonomía, la distribución y la historia natural d... more RESUMEN Se presenta nueva información sobre la taxonomía, la distribución y la historia natural de los lagartos Mabuya berengerae y M. pergravis; así como de la serpiente Coniophanes andresensis, tres especies endémicas y poco conocidas de las islas de San Andrés y Providencia, las cuales se creían muy raras e incluso extintas. A pesar del poco tiempo de estudio en las islas, se evidenció que los lagartos no son raros, encontrándose hasta ocho ejemplares cada media hora. La evaluación del estado de conservación según los parámetros de la IUCN para las tres especies, clasifica a M. berengerae y M. pergravis como especies casi amenazadas (NT) y a C. andresensis como una especie en peligro crítico de extinción (CR). La pequeña área de distribución geográfica, el impacto antrópico y la introducción de especies exóticas pueden ser las principales causas de la disminución de la población de C. andresensis. Se analizaron algunos caracteres taxonómicos en M. berengerae (hasta ahora conocida...
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Papers by José Rances Caicedo Portilla