What happened?

Description

When I set autosaveDrafts to false, toggling a checkbox or a lightswitch, with conditional fields attached to its state, inside a matrix, doesn't work : the fields that are shown don't switch, and the call to /index.php?p=admin%2Factions%2Felements%2Fupdate-field-layout&v=1730400650789 returns a 400 with this output :

{
    "name": "Bad Request",
    "message": "Invalid element UUID: 8971f694-2b94-4d7b-bb61-3f49f742d244",
    "code": 0,
    "status": 400,
    "exception": "yii\\web\\BadRequestHttpException",
    "file": "/var/www/html/vendor/craftcms/cms/src/controllers/ElementsController.php",
    "line": 2140,
    "trace": [
        {
            "file": "/var/www/html/vendor/craftcms/cms/src/controllers/ElementsController.php",
            "line": 1880,
            "function": "_element",
            "class": "craft\\controllers\\ElementsController",
            "type": "->"
        },
        {
            "function": "actionUpdateFieldLayout",
            "class": "craft\\controllers\\ElementsController",
            "type": "->"
        },
      ...
    ]
}

This happens every time when autosaveDrafts is set to false, but I've also seen it happen a few times when it's enabled, but I haven't been able to really reproduce it in a repeatable way...

Steps to reproduce

1. set autsaveDrafts to false
2. Create a matrix with an entry type that contains a lightbox and two other fields, with conditional rules between the three
3. In the CP create an entry, add the matrix, and try to toggle the switch with the devtools opened

Alternatively, here is a minimal github repo to easily reproduce it : https://github.com/ajoliveau/bug-conditional-craft
I only did :

1. ddev config --project-type=craftcms --docroot=web
2. ddev composer create -y "craftcms/craft"
3. Changed the general/config.php file to disable autosaveDrafts
4. Created a section, entry type, matrix field and the three fields

Expected behavior

Fields with conditional rendering conditions should work as intented

Actual behavior

Conditionals are not working

Craft CMS version

5.4.9

PHP version

8.2.24

Operating system and version

Linux 6.11.5-arch1-1

Database type and version

MariaDB 10.11.9

Image driver and version

Imagick 3.7.0 (ImageMagick 6.9.11-60)

Installed plugins and versions

None