Bryologie 2024 ● 45 ● 5

Directeur de la publication / Publication director: Gilles BLOCH

Président du Muséum national d’Histoire naturelle

Rédacteur en chef / Editor-in-chief: Denis LAMY

Assistant de rédaction / Assistant editor: Chris LE COQUET-LE ROUX ([email protected])

Mise en page / Page layout: Chris LE COQUET-LE ROUX

Rédacteurs associés / Associate Editors

Biologie moléculaire et phylogénie / Molecular biology and phylogeny
Bernard GOFFINET
Department of Ecology and Evolutionary Biology, University of Connecticut (United States)
Mousses d’Europe / European mosses
Isabel DRAPER
Centro de Investigación en Biodiversidad y Cambio Global (CIBC-UAM), Universidad Autónoma de Madrid (Spain)
Francisco LARA GARCÍA
Centro de Investigación en Biodiversidad y Cambio Global (CIBC-UAM), Universidad Autónoma de Madrid (Spain)
Mousses d’Afrique et d’Antarctique / African and Antarctic mosses
Rysiek OCHYRA
Laboratory of Bryology, Institute of Botany, Polish Academy of Sciences, Krakow (Pologne)
Bryophytes d’Asie / Asian bryophytes
Rui-Liang ZHU
School of Life Science, East China Normal University, Shanghai (China)
Bioindication / Biomonitoring
Franck-Olivier DENAYER
Faculté des Sciences Pharmaceutiques et Biologiques de Lille, Laboratoire de Botanique et de Cryptogamie, Lille (France)
Écologie des bryophytes / Ecology of bryophyte
Nagore GARCÍA MEDINA
Department of Biology (Botany), and Centro de Investigación en Biodiversidad y Cambio Global (CIBC-UAM), Universidad Autónoma de Madrid (Spain)

Couverture / Cover:
Photograph of Tortula revolvens turf.

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 Revisiting the Bryophyte Checklist of Algeria:
Exploring Diversity and Distribution

Rosa M. ROS
Departamento de Biología Vegetal (Botánica), Universidad de Murcia,
Campus de Espinardo, E-30100 Murcia (Spain)
[email protected] (corresponding author)

Susana RAMS
Departamento de Didáctica de las Ciencias Experimentales, Universidad de Granada,
Campus Universitario de Cartuja, E-18071 Granada (Spain)
[email protected]

Olaf WERNER
Eloy CERÓN
Departamento de Biología Vegetal (Botánica), Universidad de Murcia,
Campus de Espinardo, E-30100 Murcia (Spain)
[email protected]

Submitted on 4 October 2023 | Accepted on 29 April 2024 | Published on 10 October 2024

Ros R. M., Rams S., Werner O. & Cerón E. 2024. — Revisiting the Bryophyte Checklist of Algeria: Exploring Diversity
and Distribution. Cryptogamie, Bryologie 45 (5): 49-116. https://doi.org/10.5252/cryptogamie-bryologie2024v45a5.
http://cryptogamie.com/bryologie/45/5

ABSTRACT
A new, updated checklist of the bryophytes of Algeria was compiled from the literature. Reports were
made mainly by French botanists in the 19th and first half of the 20th centuries and include a total
of 498 taxa, of which 477 are at species level and the rest are at infraspecific rank. It comprises three
hornwort species, 118 liverwort taxa (113 species), and 377 moss taxa (361 species). For each taxon,
literature records and localities explored in the country are provided. The analysis of the current data
reveals that Algeria is the Maghreb country with the second-largest number of bryophyte taxa, be-
hind Morocco. As in other Mediterranean countries, hornworts and liverworts are scarce (24.3%),
and mosses are dominant (75.7%). Concerning liverworts and hornworts, the family Ricciaceae
and genus Riccia (31 taxa) are the best-represented; the aquatic and ephemeral genus Riella (8 taxa)
shows the higher number of species in the Mediterranean countries. Regarding mosses, the family
Pottiaceae is the largest one, with 107 taxa (28.4% of the moss taxa), and Tortula (22 taxa) is the
most species-rich genus. Algeria has, by now, the lowest species/km2 ratio (0.20 × 10-3) compared to
the rest of the Maghreb and some European Mediterranean countries (Greece, Italy, and Spain), due
KEY WORDS not only to the low degree of knowledge of the bryophyte flora but also to the larger surface area of
Northern Africa, the country and the great extension of the Saharan region. The high percentages of rare or very rare
Maghreb,
Mediterranean, taxa (72.5%) versus the low percentages of frequent, common, and very common (27.5%) can be an
mosses, indicator that many areas are still underexplored, and that the bryophyte flora of the country cannot
liverworts,
hornworts, be considered well-known. The revision or collection of new samples of some rare or doubtful taxa
updated catalogue. would be necessary to assess the reliability of some reports.

CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5) © Publications scientifiques du Muséum national d’Histoire naturelle, Paris. www.cryptogamie.com/bryologie 49
 Ros R. M. et al.

RÉSUMÉ
Révision de la liste de contrôle des bryophytes d’Algérie : exploration de la diversité et de la distribution.
Une nouvelle liste actualisée des bryophytes d’Algérie a été établie à partir de la littérature. Les récoltes
ont été faites principalement par des botanistes français au cours du 19ème et de la première moitié
du 20ème siècle et comprennent un total de 498 taxons, dont 477 sont au niveau de l’espèce et le
reste au niveau infraspécifique. Elle comprend trois espèces d’anthocérotes, 118 taxons d’hépatiques
(113 espèces) et 377 taxons de mousses (361 espèces). Pour chaque taxon, les données bibliographiques
et les localités explorées dans le pays sont fournies. L’analyse des données actuelles révèle que l’Algérie
est le pays du Maghreb qui possède le deuxième plus grand nombre de taxons de bryophytes, derrière
le Maroc. Comme dans les autres pays méditerranéens, les anthocérotes et les hépatiques sont rares
(24,3 %) et les mousses sont dominantes (75,7 %). En ce qui concerne les hépatiques et les anthocérotes,
la famille des Ricciaceae et le genre Riccia (31 taxons) sont les mieux représentés ; le genre aquatique
et éphémère Riella (8 taxons) présente le plus grand nombre d’espèces dans les pays méditerranéens.
En ce qui concerne les mousses, la famille Pottiaceae est la plus importante, avec 107 taxons (28,4 %
des taxons de mousses), et Tortula (22 taxons) est le genre qui atteint le plus grand nombre d’espèces.
L’Algérie a, à ce jour, le plus faible ratio espèces/km2 (0,20 × 10-3) par rapport au reste du Maghreb
et à certains pays méditerranéens européens (Grèce, Italie et Espagne), en raison non seulement du
MOTS CLÉS faible degré de connaissance de la flore bryophytique, mais aussi de la plus grande superficie du pays
Afrique du Nord, et de la grande extension de la région saharienne. La forte proportion de taxons rares ou très rares
Maghreb,
Méditerranée, (72,5 %) contre la faible proportion de taxons fréquents, communs et très communs (27,5 %) peut
mousses, indiquer que de nombreuses régions sont encore sous-explorées et que la flore bryophytique du pays
hépatiques,
anthocérotes, ne peut pas être considérée comme bien connue. La révision ou la collecte de nouveaux échantillons
catalogue actualisé. de certains taxons rares ou douteux serait nécessaire pour évaluer la fiabilité de certaines récoltes.

INTRODUCTION receives occurs during spring and autumn, with quantities

higher than 400 mm per year, even reaching a mean higher
Geographic characteristics of Algeria than 2000 mm per year in some points such as Kabylia
Algeria, officially the People’s Democratic Republic of and Kroumerie; the characteristic vegetation is made up of
Algeria, is in the northwest of the African continent, in Abies sp. pl., Cedrus atlantica (Endl.) Manetti ex Carrière,
the center of the Maghreb, which includes, in a restricted Olea europaea L. var. sylvestris (Mill.) Lehr, Quercus faginea
sense, Morocco, Algeria, and Tunisia. Algeria borders seven Lam., Q. ilex L., Q. suber L., and Tetraclinis articulata (Vahl)
countries: Tunisia and Libya to the east, Niger and Mali to Mast. (Charco 1999).
the south, and Mauritania, Morocco, and Western Sahara Immediately to the south of the Tellian Atlas are the Hauts
to the west. To the north, a coastline opens, more than Plateaux, in which the main steppes of northern Africa develop.
1200 km long, towards the Mediterranean Sea. The country They are high plains surrounded by mountains. Many rivers
has a surface area of 2 381 741 km2 (Oficina de Información and temporary streams flow into the plain and form lagoons
Diplomática del Ministerio de Asuntos Exteriores, Unión in the depressions that also collect surface runoff water. The
Europea y Cooperación 2023). Algeria includes two com- characteristic climate is of the extreme continental type, with
pletely different phytogeographical regions: the Mediter- very hot summers reaching 50°C and very cold winters with
ranean, to the north, where the Atlas Mountains abruptly temperatures below zero; daily thermal amplitude can reach a
shape the environment, and the Saharan-Arabian, to the variation of 40°C. Main precipitation ranges between 150 and
south (Charco 1999). The Sahara Desert covers an area of 400 mm per year, and the vegetation is made up of Q. ilex,
2 million km2 in Algeria, which represents approximately which leads to Pinus L. forests, T. articulata, Juniperus L.
85% of the territory (Oficina de Información Diplomática with needle-like leaves, Juniperus with scale-like leaves as the
del Ministerio de Asuntos Exteriores, Unión Europea y drought increases; and even to Stipa tenacissima L. formations,
Cooperación 2023). together with Artemisia sp. pl. and Rosmarinus officinalis L.
In the Mediterranean part of the country, the predominant (Charco 1999).
geographic feature is formed by the Atlas Mountains, made up Finally, in the Saharan Atlas, a long mountain range rising
of different massifs and mountain ranges; in each of them, dif- on the fringes of the Sahara Desert represents a great barrier
ferent climatic conditions and types of vegetation are present. between the Mediterranean region and the Saharan-Arabian
The massif located further north is the Tellian Atlas, situated region, with altitudes not higher than 2000 m; the climate
in the wettest area and boasting the highest concentration of is Mediterranean-steppic (dry-semi-arid) in the north slope,
forests in northern Africa, together with the Rif, in Morocco. where annual rainfall ranges from 150 to 400 mm, and
With a Mediterranean climate, 65% of the precipitation it the Mediterranean desertic (arid-semi-arid) in the south

50 CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5)

 Revisiting the Bryophyte Checklist of Algeria

slope, where mean annual rainfall does not reach 100 mm; A.-H. Letourneux (1820-1890), J. É. G. N. Paris (1827-
the vegetation corresponds to Pinus forests with Juniperus 1911) and H. Philibert (1822-1901). The French pharma-
with needle-like leaves, and with scale-like leaves (Charco cist and botanist E. Bescherelle (1828-1903) published the
1999). The Sahara Desert reaches high temperatures and collections of these botanists and other earlier ones in his
hot and dry air currents; summers are hot, and winters reach “Catalogue des Mousses observées en Algérie”, proposing
temperatures below zero in some places; in a normal year, many new taxa (Bescherelle 1882). Some of these collectors
the maximum temperature recorded is normally 48-50°C, also published their own collections (Philibert 1880a, b;
but it can reach even 56-58°C; not only are temperature Gandoger 1883).
differences recorded between summer and winter, but vari- Most of the data generated since then were studied by
ations of 20°C are also recorded between day and night, a new bryologist, L. C. Trabut (1853-1929), who lived in
which can even reach 30°C (Ozenda 1977). Precipitation is Alger until his death and made a substantial number of new
null or almost null in the Sahara, due to the great distance collections. Trabut’s work was immense; in collaboration with
that it presents to the sea; an irregular annual rainfall of J. A. Battandier (1848-1922) he carried out the inventory of
less than 50 mm has been recorded in the Central Saharan the flora of northern Africa, although only one issue of his
Region; only in the central massifs such as Hoggar and publication included five bryophytes (Battandier & Trabut
Tassili n’Ajjer, slightly more abundant and regular rains 1886). In other publications, Trabut described many species
have been observed, which even reach 200 mm per year; of mosses and liverworts and reported many new species for
no outstanding flora is found due to the harsh climatic Algeria (Trabut 1886, 1887a, b, 1888, 1908, 1913, 1914,
conditions present (Ozenda 1977). 1916, 1922, 1927, 1933, 1934; Battandier et al. 1914;
Douin & Trabut 1919). An important part of his work was
Evolution of the study of bryophytes summarized in the “Flore des Hépatiques de l’Afrique du
in Algeria over time Nord” (Trabut 1941), published after his death. The work
According to Jelenc (1955a), the first data about bryological carried out by Trabut on the study of bryophytes in northern
explorations in the Maghreb dates to 1799. They were due Africa has been, together with that of Durieu de Maison-
to the French botanist R. L. Desfontaines (1750-1833), neuve, the most important on the topic until the first half
who carried out a botanical expedition from Tunisia to of the 20th century. Some European botanists or mycologists
Algeria that lasted three years. His results were included reported his bryological data, such as A. C. Julien (x-1902)
in his work “Flora Atlantica” (Desfontaines 1799), where in the region of Constantine (Julien 1894), V. Litschauer
he mentioned 11 species of mosses and one liverwort, (1879-1939) in Alger (Litschauer 1904), and G. Lapie
nine of them collected in Algeria but without indication (fl. 1914) in the Kabylie du Djurdjura (Lapie 1908, 1909).
of their location. During this period, other botanists carried out collections
In the 19th century, botanists dedicated to the study of in Algeria, such as E. de Bergevin (1859-1933) in the whole
northern African vascular plants showed interest in bryophytes, country, R. Chudeau (1864-1921) in Constantine, the Sahara,
also collecting these plants in their expeditions. That was the and the Djurdjura, B. P. G. Hochreutiner (1873-1959) in
case of C. J. Gouget (1809-1877), who explored Algiers, the vicinity of Alger and Oran, H. Gay (1882-1909) around
Oran, and Constantine between 1835 and 1839, J.-N. Bové Blida, and P. E. Pinoy (1873-1948) in Oran and Tlemcen.
(1802-1842) who toured the Algiers region between 1837 Most of these materials were studied by J. Cardot (1860-
and 1841, and A. V. Roussel (1795-1874), who collected 1934), L. Corbière (1850-1941), C. I. Douin (1858-1944),
many species around Algiers. The last one sent the samples P. A. Hariot (1854-1917), and Trabut (Brongniart 1882; Car-
for identification to J. P. F. C. Montagne (1784-1866), who dot 1884, 1904; Corbière 1889, 1904, 1905; Douin 1904;
published the results (Montagne 1838). Camus 1906; Hariot 1913).
Between 1839 and 1882 the French government promoted R. C. J. E. Maire (1878-1949) surveyed the whole of
a scientific exploration of Algeria. It was chaired first by northern Africa and the north of the Sahara and made many
Colonel J. B. G. N. Bory de Saint-Vincent (1778-1846) and contributions, in addition to what he had done before (Maire
the botanist M. C. Durieu de Maisonneuve (1796-1878), 1916, 1936, 1937, 1939, 1940; Maire & Sevenet 1928).
with the latter serving as the primary collector of bryophytes Also, L. E. Gauthier-Lièvre (1897-1992) and J. Feldmann
due to his passion for cryptogams. (1905-1978) made some interesting additions (Gauthier-
They were in contact with Montagne, who studied most Lièvre 1931; Feldmann 1945, 1947), as well as J. Braun-
of their samples (Bory de Saint-Vincent & Montagne 1843; Blanquet (1814-1980), G. Andreánszky (1895-1967),
Durieu de Maisonneuve 1846, 1849, 1868; Montagne E. Wilczek (1867-1948) and many other botanists. Many
1849, 1856). In 1852, the French botanist and mycologist of the collections of this period were studied by M. Bizot
E. S.-C. Cosson (1819-1889) accompanied Durieu de Mai- (1905-1979), A. Coppey (1874-1913), H. N. Dixon (1861-
sonneuve on the scientific explorations, and he became the 1944), C. Meylan (1868-1941), R. A. L. Potier de la Varde
most prominent researcher of this period traveling to Algeria, (1878-1961), and I. Thériot (1859-1947). Some of the out-
but also to Tunisia and north of the Sahara (Cosson 1853). standing publications were: Dixon (1904, 1931), Coppey
Other bryophyte collectors in this period were H.-R. Le T. de (1911), Potier de la Varde (1922), Braun-Blanquet & Maire
la Perraudière (1831-1861), J. M. Gandoger (1850-1926), (1924), Thériot & Trabut (1930), Meylan (1931, 1936),

CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5) 51

 Ros R. M. et al.

Thériot (1931, 1933), Szepesfalvi (1932), and Andreán- endemic species of northern Africa, as many of them were
szky (1934). Some of them were published in the second described based on partial knowledge of the variability of
half of the 20th century, such as Dubuis & Faurel (1945), the species (Cano et al. 2000; Ros & Cano 2008; Ros et al.
Bizot & Lauriol (1950), Bizot & Potier de la Varde (1952), 2023). Between 2007 and 2013, a large group of bryolo-
Braun-Blanquet (1954), Léredde (1954), Chevassut (1956), gists working in different countries of the Mediterranean
Chevassut & Quezél (1956, 1958), Collenot et al. (1960), basin, led by R. M. Ros and V. Mazimpaka, published the
Bizot (1973), Frahm (1978). checklists of the bryophytes of the Mediterranean Basin,
Another key bryologist who deserves to be highlighted which hold the last published national catalogue of Algeria.
in the study of Algerian bryophytes is the Frenchman Ros et al. (2007a) include information on liverworts and
F. Jelenc (1911-2001), who lived in Algeria until 1962. hornworts, and Ros et al. (2013) on mosses. Both have
His herbarium in Alger was destroyed, and, unfortunately, served as a basis for this work.
many of his types disappeared (Cano et al. 2000). Only a In the last few years, some Algerian botanists at the Uni-
reduced number of his northern African collections are still versity of Badji Mokhtar (Annaba) have shown interest in
at MHLCLF herbarium (Muséum Henri Lecoq Clermont- bryophytes (Boukhatem et al. 2017). Additionally, a bryologist
Ferrand, France). group has recently emerged at the University El Bez (Setif )
He carried out numerous studies throughout northern Africa and initiated studies in the Setif region (Mazari & Boulaacheb
and published many bryological papers (Jelenc 1948, 1949, 2023; Mazari et al. 2024).
1950, 1951, 1953, 1954a, b, 1955a, b, 1956, 1957, 1959, The main aim of this paper is to compile published infor-
1966, 1967). Their results were compiled in the work entitled mation about Algerian bryophytes, including literature refer-
“Muscinées de l’Afrique du Nord (Algérie, Tunisie, Maroc, ences and localities where species were reported. That should
Sahara)” (Jelenc 1955a). This is an exhaustive inventory of be the basis for a modern and updated Algerian bryophyte
the citations already published and their unpublished data. flora. The specific objectives are: 1) to know the state of the
The work holds a catalogue of 159 liverworts and hornworts art of Algerian bryology that should be kept in mind for new
and 444 mosses, with an indication of the localities grouped Algerian studies; 2) to state which Algerian areas have been
by geographical areas. A few years later, he published a sup- sampled in the past and where new floristic studies should be
plement (Jelenc 1967) with the results of his later studies, undertaken; 3) to draw attention to species in the catalogue
which was his last publication. that should be reviewed to establish a trustworthy list of the
It is remarkable that two women, the French hepaticolo- Algerian bryophytes; and 4) to lay the groundwork for the
gists S. Jovet (1914-2006) and H. Bischler (1932-2005) protection of Algerian bryophytes.
worked very intensively in northern Africa and all over the
Mediterranean basin. Unfortunately, they never published
a catalogue of liverworts from Algeria. Nevertheless, they MATERIAL AND METHODS
mentioned their presence in the country when giving distri-
bution data in other publications of the Mediterranean Basin Criteria for the list
(Jovet-Ast 1955, 1956; Jovet-Ast & Bischler 1968, 1971) All the publications known to contain any reference to Algerian
and taxonomical studies (Bischler 1970, 1978). Their most bryophytes were compiled, and the reports were introduced
outstanding works were the taxonomic revision of the genus in a database according to the original names used in each
Riccia in the Mediterranean Region by Jovet-Ast (1986) and publication. The checklist therefore includes all the published
the book about the Mediterranean liverworts by Bischler known records of Algerian bryophytes. Unlike what was done
(2004) which hold vast information about the presence of in the previous Algerian checklists published by Ros et al.
this group of bryophytes in Algeria. (1999, 2007a, 2013), which were based on the compilations
The rest of the works published in the second half of the of Jelenc (1955a, 1967), here all known published records
20th century have very little original data. They are mostly have been included from 1799 (Desfontaines 1799) up to
punctual revisions of previously published data. In the last the present day.
two decades of the 20th century, the northern African bryo- The veracity of the reports has not been revised. This paper
phytes attracted the interest of Spanish bryologists due to is a compilation of the published data to ease the necessary
their proximity and the similarity of their floras. Research- posterior work for a critically revised checklist.
ers from the University of Murcia started a research line The European checklist by Hodgetts et al. (2020) was the
on the Mediterranean Bryophyte Flora and made several main basic reference for accepted names and taxonomic
collecting trips to Algeria (Ros et al. 1990). They reviewed adscription of genera to families, as most of the reported
Jelenc’s checklists to assess the breadth of their discoveries taxa are also present in Europe. Some exceptions were
and published a new bryophyte checklist of northern Africa made in the light of more recently published taxonomic
(Ros et al. 1999) using updated taxonomical and nomen- papers or other criteria, and these are documented after the
clatural criteria. In addition to Algeria, other Maghreb accepted names, such as those related to the genera Cal-
countries were considered, such as Morocco and Tunisia, cidicranella Bonfim Santos, Fedosov & Jan Kučera (Fedosov
but also Egypt, Libya, Mauritania, Western Sahara, the et al. 2023), Metaneckera Steere (Enroth et al. 2022) and
north of Niger, and Mali. In a parallel way, they studied the Oxystegus (Limpr.) Hilp. (Zander & Eckel 2019), and the

52 CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5)

 Revisiting the Bryophyte Checklist of Algeria

species Microbryum davallianum (Sm.) R.H.Zander (Ros For the name of a given locality, which can vary between
et al. 1996), Neotrichostomum crispulum (Bruch) R.H.Zander different publications, in general, the spelling of the first
(Zander 2023), and Weissia armata (Thér. & Trab.) Fedosov author who cited the taxon is used. When the name of the
(Fedosov 2011). locality is confusing and does not coincide with the form
When accepted names are different from those used in the used by other authors, the name used by the first author is
last Algerian checklists (Ros et al. 2007a, 2013), the names followed in parentheses by the form used by other authors,
used in these works are given to help the identification of e.g. Lac Touga (Tonga). When the localities were imprecise
taxa and comparison between former and present checklists. over a large area and there was also published information
For synonyms, Ros et al. (2007a, 2013) are the basic refer- about more precise localities that belong to this large area,
ences, but when newly proposed ones and older ones are not the imprecise locality was also left, since sometimes this
included in any of them, they are given here after the cur- indicates that the species is widespread throughout that
rently accepted name, indicating the work where the syno- area (e.g. Monts et Haute-Plaine de Tlemcem and Haute
nyms were published. In cases where infraspecific taxa were Plaine de Tlemcen, Berges du Chabet bel Abbès, 750 m).
not distinguished in the literature, the relevant occurrences Finally, the abundance in Algeria of each accepted taxon
are shown in the checklist at the species level (e.g. Fissidens is indicated.
bryoides instead of F. bryoides var. bryoides).
Taxa names are arranged alphabetically by genus and species. Flora analysis
Two groups have been considered, hornworts and liverworts Analysis of species diversity, species richness, and species
on one side, and mosses on the other side. abundance were made. For species diversity, the number of
Accepted names are written in both italics and bold. Taxa taxa of the three bryophyte groups (liverworts, hornworts,
for which there are doubts about their taxonomical validity, as and mosses) was analysed, as well as their biotypes: thal-
well as the invalid names, are shown only in italics. All entries lose versus foliose in the two first, and acrocarpous versus
are also arranged alphabetically in the list, with indications pleurocarpous in the mosses, as they have different degrees
of literature reports and localities, along with explanations or of adaptation to Mediterranean conditions. Also, they were
justifications of their current status. examined at family and genus levels to determine the more
Additionally, species currently accepted by the scientific diverse taxonomical groups.
community and listed in Hodgetts et al. (2020), but facing For the species richness analysis, a comparison was made
doubts regarding their presence in Algeria, are italicized and between Algeria and the other two Maghreb countries (in
marked with an asterisk (*). These doubts may stem from the restricted sense, Morocco and Tunisia), as well as Algeria
changes in taxonomic criteria used to define the species since and three European Mediterranean countries with different
the publication of the record, errors in synonymy utilized extensions and depths of bryological exploration (Greece, Italy,
in previous works, the lack of evidence listing the species and Spain). As the whole of Algeria belongs to continental
in Algeria, or identification uncertainties as assessed by the Africa, only the continental parts of these countries were
authors of this work. For these last cases, the revision of the considered; therefore, islands and archipelagos were excluded.
original material would be desirable. However, the high The number of taxa in each country was taken from Ros et al.
dispersion of herbarium collections, which were often made (2007a, 2013) for liverworts and hornworts on the one hand
by non-bryologists and sent for identification to specialists, and mosses on the other hand, except for Italy, for which the
poses challenges in locating the samples. Moreover, the disap- more recent checklist of Aleffi et al. (2020) was used. To know
pearance of a significant portion of Jelenc’s herbarium, one the bryophyte number present in 1 km2 of each country, the
of the most important collections for Algerian bryophytes, taxa numbers were related to the total area of each country,
further complicates the assessment of the identity of many which were obtained from the Geography of Greece (2022)
doubtful taxa in the future. Explanations are provided for and IberiaNature (2022) for the Greek and Spanish mainland
each case. Literature reports based on the revision of her- surfaces, respectively, and the United Nations (2017) for the
barium material, despite no locality being indicated (sine Italian surface, but modified according to Sardegna (2022)
loco) are accepted as correct. The work of Bischler (2004), and Sicilia (2022).
related to liverworts and hornworts of the Mediterranean, The species abundance of each accepted taxon in Algeria
which explicitly indicates that specimens were collected in was estimated from the scale proposed by Huarte Irurzun
Algeria but that localities were only available on CD upon (2001), later used by Rams et al. (2014), but with modi-
request, deserves a special mention. Unfortunately, the CD fications to adapt it to this study, as the highest number of
seems to have been lost (Denis Lamy, personal communica- localities for a given taxon in Algeria was 55, much lower than
tion), and Bischler’s Algerian localities are unknown to the in the two mentioned studies that were near 100. The scale
authors of this work. Taxa excluded from Algeria are also used is the following: very rare (present only in one locality),
written in italics and preceded by two asterisks (**). All the rare (present in 2-10 localities), frequent (present in 11-20
published literature sources containing Algerian records are localities), common (present in 21-40 localities), very com-
given for each taxon, arranged chronologically. They are mon (present in more than 40 localities). Reports “sine loco”
grouped by localities to highlight the areas and geographi- have not been considered for abundance unless they are the
cal areas surveyed. only ones accepted for a given species.

CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5) 53

 Ros R. M. et al.

RESULTS Asterella africana (Mont.) A.Evans

Records. — Montagne (1838): Alger. Litschauer (1904), Jelenc
Liverworts and hornworts (1955a): Col de Tirourda, 1700 m. Montagne (1856), Trabut (1941),
Bischler (2004): sine loco. Rare.
Aneura pinguis (L.) Dumort.
Calypogeia arguta Nees & Mont.
Records. — Montagne (1838): Alger. Jelenc (1955a): Atlas de
Blida; Kabylie du Djurdjura, Ait Ouaban. Chevassut & Quézel Records. — Bischler (1970): sine loco. Rare.
(1956): La Réghaïa. Jelenc (1967): Kabylie du Djurdjura, Ak-
fadou. Rare.
Comment
This author based the presence of the species in Algeria on the
*Anthoceros agrestis Paton study of three specimens from different herbaria (P, S-PA, W).

Comment
Calypogeia azurea Stotler & Crotz
Doubtfully present in Algeria. Ros et al. (2007a) included it
in the Algerian list as a “Report without precise collection Records. — Trabut (1941), Jelenc (1955a): Bône; Edough. Feld-
data or locality”, based erroneously on Jelenc (1955a), who mann (1947), Jelenc (1955a): Kabylie de Collo, vallée de l’Oued Bin
included its synonym A. crispulus (Mont.) Douin as sine loco el Ouidante, près de Taoulel, 600 m. Jelenc (1954b, 1966, 1967):
Kabylie du Djurdjura dans le Massif de l’Akfadou. Jelenc (1955a):
for northern Africa based on Trabut information, but not Aumale, Oued Hamidou. Jelenc (1959): Kabylie de Collo, Baie de
for Algeria. It has often been confused with A. punctatus in Tamamart, 50 m. Rare.
old literature (Söderström et al. 2002), which led Ros et al.
(2007a) to think that most of the Mediterranean reports Comment
should belong to A. punctatus. Following Söderström et al. (2002), reports in Algeria refer-
ring to the name C. trichomanis auct. have been attributed
to C. azurea. However, considering that the concept of
Anthoceros punctatus L. C. trichomanis often encompasses C. muelleriana and poten-
tially other species, the presence and distribution in Algeria
Records. — Litschauer (1904), Jelenc (1955a): Fort National, of this complex of species is doubtful.
1000 m. Trabut (1941): sine loco. Feldmann (1947), Jelenc (1955a):
Kabylie de Collo, Baie de Tamanart; Kabylie de Collo, vallée de
l’Oued Elli Ankou, 500-600 m. Jelenc (1955a): Souk Ahras. Jelenc
(1955a), Ahayoun et al. (2007): Alger. Rare. Calypogeia fissa (L.) Raddi
Records. — Trabut (1941): Oued-el-Akhal. Bischler (1970): Djurd-
jura, Akfadou; Edough. Bischler (2004): sine loco. Rare.
Apopellia endiviifolia (Dicks.) Nebel & D.Quandt
*Calypogeia muelleriana (Schiffn.) Müll. Frib.
Pellia endiviifolia (Dicks.) Dumort.

Records. — Corbière (1904): Tlemcen, entre Sidi bou Médine Comment

et les cascades. Corbière (1904), Jelenc (1955a): Sidi-Bel-Abbès. Doubtfully present in Algeria. Bischler (2004) reported that
Braun-Blanquet & Maire (1924), Trabut (1941), Jelenc (1955a): it had been recorded previously in Algeria, but no published
Tlemcen. Trabut (1941), Jelenc (1955a): Alger; Tiaret. Jelenc (1949): reports have been found under this name. It could be based on
Dans les monts et la haute plaine de Tlemcen. Jelenc (1951): Monts
de Tlemcen, cirque d’el Ourit. Jelenc (1951, 1967): Haute plaine de
some old C. trichomanis reports, as Bischler (1970) informed
Tlemcen, cascades de l’oued Bou-Ennag à la traversée des collines that some African records she revised belong to C. muelleriana,
des Beni Mester, 500 m. Jelenc (1953): Aurès, flanc nord du Ras although no precise information was published.
Pharaoun, dans les torrents prés d’Aïn Mimoun. Jelenc (1953, 1955a):
Aurès, Ravin de la maison forestére du Chélia, dans les cascades.
Jelenc (1956): Sahel d’Alger, dans le ravin du Frais-Vallon. Jelenc
(1966, 1967): Kabylie du Djurdjura dans la Massif de l’Akfadou. Cephalozia bicuspidata (L.) Dumort.
Bischler (2004): sine loco. Frequent.
Records. — Corbière (1889), Jelenc (1955a): Blida, près la glacière
Laval. Bischler (2004): sine loco. Very rare.
Comment
Pellia endiviifolia fo. undulata Jelenc nom. inval. was reported Cephaloziella baumgartneri Schiffn.
from Monts de Tlemcen, El Ourit by Jelenc (1951, 1967) but
according to Ros et al. (2007a) no other record or reference Records. — Trabut (1941), Jelenc (1955a): Alger; Blida; Mosta-
to it has been found. ganem. Jelenc (1955a): La Bouzaréa. Ros et al. (1990): Between

54 CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5)

 Revisiting the Bryophyte Checklist of Algeria

Mostaganem and Cap Ivi, Abdelmaimalek-Ramdan beach, 0 m; Comment

Hammam-Riga, 500 m. Bischler (2004): sine loco. Rare. Ros et al. (2007a) did not distinguish var. rivularis within
Chiloscyphus polyanthos, therefore the reports of this taxon
were joined with the typical variety.
Cephaloziella calyculata (Durieu & Mont.) Müll. Frib.

Records. — Durieu de Maisonneuve (1849), Stephani (1889a), Clevea hyalina (Sommerf.) Lindb.
Bischler (2004): sine loco. Montagne (1856), Trabut (1941), Jel-
enc (1955a): Prope Philippeville. Corbière (1904): Mostaganem, Athalamia hyalina (Sommerf.) S.Hatt.
120 m. Jelenc (1955a, 1966, 1967): Alger au Fort des Arcades.
Jelenc (1955a): Oran. Rare. Nomenclature follows Rubasinghe et al. (2011) and Söderström
et al. (2016).
Records. — Trabut (1914, 1941), Jelenc (1955a): Djurdjura, Forêt
Cephaloziella divaricata (Sm.) Schiffn. des Aït-Ouaban. Trabut (1941), Jelenc (1955a): Gertoufa près Ti-
aret; Mazer, cascades de l’Oued Khemis. Jelenc (1955a): Kabylie du
Records. — Corbière (1904), Jelenc (1955a): La Bouzaréah, près Djurdjura, Tirourda, 1800 m; Sersou, Guertoufa. Bischler (2004):
Alger. Corbière (1905), Jelenc (1955a): Environs de Constantine, sine loco. Rare.
Djebel Ouach. Trabut (1941): Alger. Jelenc (1955a): El Biar. Bischler
(2004): sine loco. Rare.
Clevea spathysii (Lindenb.) Müll.Frib.
Cephaloziella stellulifera (Taylor ex Spruce) Schiffn. Athalamia spathysii (Lindenb.) S.Hatt.

Records. — Trabut (1941), Jelenc (1955a): Constantine; Forêt Nomenclature follows Rubasinghe et al. (2011) and Söderström
de la Réghaia. Jelenc (1967), Bischler (2004): sine loco. Ros et al. et al. (2016).
(1990): Texenna’s wood, 700 m. Rare. Records. — Montagne (1838), Jelenc (1955a): Alger. Montagne
(1856), Bischler (2004): sine loco. Corbière (1904): Alger-Mustapha,
120 m. Trabut (1941): les environs d’Alger. Jelenc (1955a): Blida;
Cephaloziella turneri (Hook.) Müll. Frib. Oran, Ravins de Noiseaux. Jelenc (1967): Tiaret, 1000 m. Rare.

Records. — Feldmann (1947), Jelenc (1955a): Kabylie de Collo,

les crêtes de Boudra, 500-600 m; Kabylie de Collo, Taoulel. Bischler Cololejeunea rossettiana (C.Massal.) Schiffn.
(2004): sine loco. Rare.
Records. — Dismier (1930), Jelenc (1955a): Gorges de la Chiffa,
ruisseau des Singes. Very rare.
Chiloscyphus pallescens (Ehrh. ex Hoffm.) Dumort.
var. fragilis (Roth) Müll. Frib.
Conocephalum conicum (L.) Dumort.
Nomenclature follows Söderström et al. (2016).
Records. — Brongniart (1882): Blidah. Corbière (1889), Jelenc
Records. — Feldmann (1947), Jelenc (1955a): Kabylie de Collo, (1955a): Blida, sources du Bou-Rebbou. Corbière (1904), Jelenc
Oued Bin el Ouidane près de Taoulel. Rare. (1955a): Bône, flancs de l’Edough. Trabut (1941), Bischler (2004):
sine loco. Jelenc (1955a): Alger; Kabylie du Djurdjura, Tiroual. Rare.
Comment Comment
Only this variety of C. pallescens has been reported in Algeria, As predicted by Ros et al. (2007a), C. salebrosum Szweyk.,
which was considered by Ros et al. (2007a) as a synonym of Buczkowska & Odrzykoski, a species split from C. conicum s.s.
var. pallescens. by Szweykowski et al. (2005), is currently widespread in the
Mediterranean area (Hallingbäck 2019). Therefore, its pres-
ence in Algeria is probable, and the distribution of conicum
Chiloscyphus polyanthos (L.) Corda var. polyanthos is uncertain.

Records. — Jelenc (1955a): Kabylie du Djurdjura, Afkadou.

Very rare.
Corsinia coriandrina (Spreng.) Lindb.
Records. — Corbière (1904), Trabut (1941): Ravin de Telemly près
Chiloscyphus polyanthos var. rivularis Alger. Corbière (1905), Trabut (1941), Jelenc (1955a): Environs de
(Schrad.) Lindb. & Arnell Constantine, Djebel Ouach. Trabut (1914, 1941), Jelenc (1955a):
Forêt d’Akfadou. Trabut (1941), Jelenc (1955a): Tiaret; Tlemcen.
Records. — Trabut (1941), Jelenc (1955a): La Calle. Jelenc Trabut (1941), Jelenc (1955a), Chevassut & Quézel (1956): Réghaïa.
(1955a): Kabylie du Djurdjura, Fontaine de Tala Semda, col de Chevassut & Quézel (1956): La Rassauta. Jelenc (1956, 1967): Sahel
Afkadou. Rare. d’Alger. Bischler (2004): sine loco. Rare.

CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5) 55

 Ros R. M. et al.

Diplophyllum albicans (L.) Dumort. Fossombronia mittenii Tind.

Records. — Bischler (2004): sine loco. Very rare. Records. — Trabut (1941), Jelenc (1955a): Alger; Réghaia. Rare.

*Dumortiera hirsuta (Sw.) Nees Fossombronia pusilla (L.) Nees

Comment Records. — Corbière (1889), Jelenc (1955a): Blida. Litschauer

(1904), Jelenc (1955a): Michelet, 1000 m. Jelenc (1955a): Alger;
Doubtfully present in Algeria. Bischler (2004) informed that El Affroun; La Bouzaréa; Marengo. Bischler (2004): sine loco. Rare.
it had been recorded previously in Algeria, but no published
reports have been found.
Fossombronia wondraczekii (Corda) Lindb.

Exormotheca welwitschii Steph. Records. — Montagne (1838), Corbière (1904), Trabut (1941),
Jelenc (1955a): Alger. Corbière (1904), Trabut (1941), Jelenc (1955a):
La Bouzaréah, près Alger, 220 m. Chevassut & Quézel (1956): Forêt
Records. — Douin & Trabut (1919), Trabut (1941), Ahayoun de Pins d’Alep de Saint-Ferdinand; La Réghaïa. Rare.
et al. (2007): Tiaret, Vallon de Takdempt à Gertonfa. Jelenc (1955a):
Sersou, Tagdempt. Bischler (2004): sine loco. Rare.
Frullania dilatata (L.) Dumort.
Fossombronia angulosa (Dicks.) Raddi
Records. — Corbière (1889), Jelenc (1955a): Blida. Corbière
Records. — Corbière (1889), Jelenc (1955a): Blida. Corbière (1904), Jelenc (1955a): La Bouzaréah, près Alger, 200 m. Corbière
(1904): La Bouzaréah, près Alger, 110 m. Corbière (1904), Jelenc (1904), Jelenc (1955a): La Calle, Forêt de l’Oued Mellah. Corbière
(1955a): Mostaganem, 198 m. Lapie (1909), Jelenc (1955a): Kabylie (1905), Jelenc (1955a): Environs de Constantine, Djebel Ouach;
du Djurdjura, Fort National. Trabut (1941), Jelenc (1955a): Alger; Environs de Constantine, Menidj. Jelenc (1955a): Djidjelli; Edough;
Bône; Edough; Tlemcen. Bischler (2004): sine loco. Rare. Kabylie du Djurdjura, Forêt d’Afkadou. Jelenc (1955a), Ahayoun
et al. (2007): Gorges de Keddara. Bischler (2004): sine loco. Rare.

Fossombronia caespitiformis De Not. ex Rabenh.

subsp. caespitiformis Frullania tamarisci (L.) Dumort.

Records. — Trabut (1887a): D’Oran a Mécheria. Corbière (1889), Records. — Feldmann (1947), Jelenc (1955a, 1959): Kabylie de
Jelenc (1955a): Blida. Corbière (1904): Alger, chemin de Telemly, Collo, Baie de Tamanart, moins de 50 m. Jelenc (1955a): Atlas de
80 m. Corbière (1904), Jelenc (1955a): La Bouzaréah, près Alger, Blida. Rare.
230 m; Le Corso; Orléansville, bois de la Pépinière. Corbière (1905),
Jelenc (1955a): Environs de Constantine, Route de Sétif. Trabut
(1941), Bischler (2004): sine loco. Jelenc (1954b): Hydra, 200 m; *Fuscocephaloziopsis connivens
Sahel d’Alger: colline du Hamma, près du Musée. Jelenc (1955a): (Dicks.) Váňa & L.Söderstr.
Alger, Frais Vallon; Castiglione; El Affroun; Littoral Oranais. Che-
vassut & Quézel (1956): La Rassauta. Chevassut & Quézel (1956), Cephalozia connivens (Dicks.) Lindb.
Jelenc (1966): Forêt de Pins d’Alep de Saint-Ferdinand; La Réghaïa.
Ros et al. (1990): Azazga; Between Mostaganem and Cap Ivi, Ab- Nomenclature follows Söderström et al. (2016).
delmeimalek-Ramdan beach, 0 m; Hammam-Riga, 500 m. Frequent.
Comment
Doubtfully present in Algeria. Bischler (2004) informed that
Fossombronia caespitiformis subsp. multispira it had been recorded previously in Algeria, but no published
(Schiffn.) J.R.Bray & D.C.Cargill in Stotler et al. reports have been found.

Records. — Trabut (1941), Jelenc (1955a): Alger; Blida; Mouzaia.

Jelenc (1955a): Cap Matifou; Castiglione; El Biar; Marengo. Che-
vassut & Quézel (1956): La Réghaïa. Chevassut & Quézel (1956),
*Fuscocephaloziopsis lunulifolia
Jelenc (1966): Forêt de Pins d’Alep de Saint-Ferdinand. Rare. (Dumort.) Váňa & L.Söderstr.

Cephalozia lunulifolia (Dumort.) Dumort.

Fossombronia echinata Macvicar Nomenclature follows Söderström et al. (2016).
Records. — Macvicar (1911), Jelenc (1955a): Belida, Mouzaiaville. Comment
Trabut (1941), Jelenc (1955a): Alger; Blida; Oran. Jelenc (1949):
Monts de Tlemcen: forêt de Zariffet, 1100 m. Chevassut & Quézel Doubtfully present in Algeria. Bischler (2004) informed that
(1956): Forêt de Pins d’Alep de Saint-Ferdinand. Ros et al. (1990): it had been recorded previously in Algeria, but no published
Djebel Chenoua, 5-40 m. Bischler (2004): sine loco. Rare. reports have been found.

56 CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5)

 Revisiting the Bryophyte Checklist of Algeria

Gongylanthus ericetorum (Raddi) Nees Lophocolea heterophylla (Schrad.) Dumort.

Records. — Philibert (1882), Trabut (1941), Jelenc (1955a): Records. — Bischler (2004): sine loco. Very rare.
Bône. Trabut (1941), Jelenc (1955a): Alger. Jelenc (1951, 1967):
Monts de Tlemcen, Forêt de Zariffet, 1200 m. Jelenc (1955a): La
Réghaia. Jelenc (1967): Kabylie du Djurdjura, Gorges de Palestro,
500 m. Ros et al. (1990): Texenna’s wood, 700 m. Bischler (2004): Lophocolea minor Nees
sine loco. Rare.
Records. — Bischler (2004): sine loco. Very rare.

Gymnocolea inflata (Huds.) Dumort.

Lunularia cruciata (L.) Lindb.
Records. — Trabut (1941): La Calle. Very rare.
Records. — Desfontaines (1799), Trabut (1941), Bischler (2004):
sine loco: Montagne (1838), Jelenc (1955a): Alger. Trabut (1887a):
D’Oran a Mécheria. Corbière (1889): Blida. Corbière (1904),
Isopaches bicrenatus (Schmidel ex Hoffm.) H.Buch Jelenc (1955a): Le Corso; Orléansville. Julien (1894): Région de
Constantine. Litschauer (1904), Jelenc (1955a): Fort National,
Lophozia bicrenata (Schmidel ex Hoffm.) Dumort. 1000 m; Qued-Kebir bei Blidah, 200 m. Corbière (1905), Jelenc
(1955a): Environs de Constantine, Djebel Ouach. Braun-Blan-
Nomenclature follows Söderström et al. (2016). quet & Maire (1924), Jelenc (1955a): Tlemcen. Trabut (1941):
Djurdjura, 2000 m. Jelenc (1949): Djebel Murdjadjo (Oran),
Records. — Corbière (1889), Jelenc (1955a): Blida, près la glacière Ravin de la Vierge; El Kalaa (Tlemcen) vers 800 m; Vallée de
Laval. Very rare. l’Isser “El Kasbat”, 500 m. Jelenc (1949, 1967): Monts et haute
plaine de Tlemcen. Jelenc (1955a): Ait Ouabane; Alma. Chevas-
sut & Quézel (1956): Forêt de Pins d’Alep de Saint-Ferdinand;
Jungermannia atrovirens Dumort. La Réghaïa. Jelenc (1956): Kabylie du Djurdjuria le massif de
l’Akfadou. Jelenc (1956, 1967): Atlas Mitidijien, pic Mouzaïa.
Jelenc (1967): La Réghaia. Ros et al. (1990): Texenna’s wood,
Records. — Trabut (1914, 1941), Jelenc (1955a): Djurdjura. 700 m. Common.
Very rare.

*Jungermannia pumila With. Mannia androgyna (L.) A.Evans

Records. — Corbière (1889), Jelenc (1955a): Blida, l’oued Si-

Comment di-el-Kebir. Corbière (1904), Jelenc (1955a): Ravin d’El Biar près
Doubtfully present in Algeria. Bischler (2004) informed that Alger, 100 m. Litschauer (1904), Jelenc (1955a): Michelet, 1000 m.
it had been recorded previously in Algeria, but no published Lapie (1909), Jelenc (1955a): Kabylie du Djurdjura, Fort National.
reports have been found. Trabut (1941), Bischler (2004): sine loco. Jelenc (1955a): Alger; La
Bouzaréa. Rare.

Lejeunea cavifolia (Ehrh.) Lindb.

Mannia atlantica Jelenc nom. inval.
Records. — Corbière (1889): Blida, derrière Mimich. Corbière
(1904), Trabut (1941): Gorges de la Chiffa, ruisseau des Singes Grimaldia dichotoma var. atlantica Trab. nom. nud.
Kabylie. Trabut (1941), Jelenc (1955a): Akfadou. Feldmann
(1947), Jelenc (1955a, 1959): Kabylie de Collo, Baie de Tamanart, Grimaldia atlantica Trab. nom. inval.
moins de 50 m. Jelenc (1955a): Gorges de la Chiffa, ruisseau des Records. — Trabut (1914): Djurdjura. Trabut (1941), Jelenc
Singes; Kabylie du Djurdjura, Djebel Afroun. Bischler (2004): (1955a): Atlas, Fort National. Jelenc (1955a): Gorges de la Chiffa.
sine loco. Rare.

Comment Comment
These seem to be the only references to these names (Ros et al.
The reports of the name Lejeunea serpyllifolia Lib. by Corbière
2007a) and their identity seems to be unresolved.
(1889, 1904) have been adscribed to L. cavifolia, following
the criterion of Ros et al. (2007a).
Marchantia paleacea Bertol.
Lophocolea bidentata (L.) Dumort. Records. — Trabut (1914), Jelenc (1955a): Djurdjura. Trabut
(1941): Fontaine à l’entrée des gorges du Chabet, route de Bougie
Records. — Trabut (1941), Jelenc (1955a): La Calle. Feldmann à Sétif. Jelenc (1949, 1967): Haute plaine de Tlemcen, berges de
(1947), Jelenc (1955a): Kabylie de Collo, Oued Medjerrah près de l’Oued Saf-Saf entre 600-650 m (entre le Pont de Mascara et El
Bou Noghra; Kabylie de Collo, Oued Melkach. Rare. Ourit). Jelenc (1955a): Kabylie des Babors, Chabet el Akra. Rare.

CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5) 57

 Ros R. M. et al.

Marchantia polymorpha L. species that grows on flushed rock faces or wet peaty ground
(Blockeel et al. 2014a) in Algeria is here rejected.
Records. — Gandoger (1883): sine loco. Feldmann (1947) Jelenc
(1955a): Kabylie de Collo, Oued Elli Ankou, 500-600 m; Kabylie
de Collo, Oued Medjerrah, 500-600 m. Rare.
Myriocoleopsis minutissima
(Sm.) R.L.Zhu, Y.Yu & Pócs
Comment
No subspecies have been distinguished in the Algerian lit- Cololejeunea minutissima (Sm.) Schiffn.
erature; therefore, the reports should refer to Marchantia Nomenclature follows Söderström et al. (2016).
polymorpha s.l.
Records. — Trabut (1941), Jelenc (1955a): Forêts de la Calle.
Bischler (2004): sine loco. Very rare.
Marsupella emarginata (Ehrh.) Dumort.
Nardia scalaris Gray
Records. — Bischler (2004): sine loco. Very rare.
Records. — Trabut (1888): sine loco. Trabut (1941), Jelenc (1955a):
Réghaia. Very rare.
Mesoptychia collaris (Nees) L.Söderstr. & Váňa
Oxymitra incrassata (Brot.) Sérgio & Sim-Sim
Leiocolea collaris (Nees) Schljakov
Nomenclature follows Söderström et al. (2016). Records. — Montagne (1838): Alger. Corbière (1904): Mostaganem,
170 m. Corbière (1904), Jelenc (1955a): La Bouzaréah, près Alger,
Records. — Bischler (2004): sine loco. Very rare. 220 m. Corbière (1905), Jelenc (1955a): Environs de Constantine,
Bekcina, 650 m; Environs de Constantine, Djebel Ouach. Trabut
(1941), Bischler (2004): sine loco. Jelenc (1949, (1967): Haute plai-
ne de Tlemcen, colline de Bréa, 600 m. Jelenc (1953): Babor, flanc
Mesoptychia turbinata (Raddi) L.Söderstr. & Váňa nord du Tababor, 1700 m. Jelenc (1955a): Kabylie du Djurdjura,
Akfadou; Sidi Daoud. Jelenc (1955a), Chevassut & Quézel (1956):
Leiocolea turbinata (Raddi) H.Buch La Réghaia. Chevassut & Quézel (1956): Forêt de Pin d’Alep de
Bäinem; Forêt de Pin d’Alep de Bäinem; Forêt de Pin d’Alep d’El
Nomenclature follows Söderström et al. (2016). Riath; La Rassauta. Jelenc (1956): Atlas mitidjien, au barrage du
Hamiz; Massif de la Bouzarèa, Forêt de Baïnem. Ros et al. (1990):
Records. — Corbière (1905), Jelenc (1955a): Environs de Con- Oued Rhiou. Frequent.
stantine, Chettabah; Environs de Constantine, Menidj; Environs
de Constantine, les Pins; Environs de Constantine, Route de la
Corniche; Environs de Constantine, Sidi M’Cid. Trabut (1941),
Bischler (2004): sine loco. Jelenc (1951): Monts de Tlemcen, cirque Pedinophyllum interruptum (Nees) Kaal.
d’el Ourit. Jelenc (1955a): Cherchell; Kabylie du Djurdjura, Ait
Abd el Ali, 900 m; La Bouzaréa Jelenc (1956): Kabylie du Djurd- Records. — Lapie (1909), Dubuis & Faurel (1945), Jelenc (1953,
jura, dans les gorges de Palestro, vers 500 m près de Hamman 1955a, 1967): Djurdjura, Plateau de l’Aizer, 1900 m. Very rare.
Tellat. Rare.

Pellia epiphylla (L.) Corda

Metzgeria furcata (L.) Dumort.
Records. — Corbière (1904): Bône, flancs d’Edough. Trabut
Records. — Feldmann (1947), Jelenc (1955a): Kabylie de Collo, (1941): Aïn-Taya. Lapie (1909): Kabylie du Djurdjura. Trabut
Oued Elli Ankou, 600 m. Very rare. (1941), Jelenc (1955a): Bône; Edough. Feldmann (1947), Jelenc
(1955a): Kabylie de Collo, vallée de l’Oued Ramèche, 600-800 m.
Jelenc (1955a): Alger; Atlas de Blida; Kabylie du Djurdjura, Dra
**Moerckia hibernica (Hook.) Gottsche Inguel; La Bouzaréa. Jelenc (1956): Kabylie du Djurdjura, Mas-
sif de l’Akafadou à la Fontaine des Ifs. Bischler (2004): sine loco.
Comment Frequent.
Excluded from Algeria. Trabut (1941) reported that there
exists a collection supposedly collected in the “cascades de Petalophyllum ralfsii (Wilson) Nees & Gottsche
Tlemcen” by Dr. Zoernig and identified by Stephani, but
that after having searched the cascades himself, he only found Records. — Battandier & Trabut (1886), Trabut (1887b): Circa
Apopellia endiviifolia (sub Pellia fabroniana). He also com- Alger, Hydra; Circa Alger, Mustapha. Battandier & Trabut (1886),
mented that the thallus of M. hibernica closely resembles Trabut (1887b), Trabut (1941), Jelenc (1955a): Circa Alger, Si-
that of P. endiviifolia. He, therefore, considers his presence in di-Ferruch. Corbière (1904), Trabut (1941), Jelenc (1955a): Or-
léansville, bois de la Pépinière, 140 m. Trabut (1941): Birmandreis.
Algeria doubtful. The report was compiled by Jelenc (1955a), Jelenc (1949): Massif du Murdjadjo, versant nord du col de Santa
indicating the locality “Monts de Tlemcem, cascades d’El Cruz. Jelenc (1955a): Sidi Merzoug. Jelenc (1967): Oran. Bischler
Ourit”. The presence of this Circumpolar Boreo-temperate (2004): sine loco. Rare.

58 CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5)

 Revisiting the Bryophyte Checklist of Algeria

*Phaeoceros carolinianus (Michx.) Prosk. Porella cordaeana (Huebener) Moore

Comment Records. — Trabut (1941), Jelenc (1955a): Kabylie; La Calle.

Jelenc (1953, 1967): Ouarsenis, cédraie de Téniet el Had, 1500 m.
Doubtfully present in Algeria. Jovet-Ast & Bischler (1968) Jelenc (1955a): Djurdjura, 2000 m; Kabylie du Djurdjura, Akfadou;
informed that it had been recorded previously in Algeria, but Kabylie du Djurdjura, Tala Ghizan. Rare.
no published reports have been found.
Porella obtusata (Taylor) Trevis.
Phaeoceros laevis (L.) Prosk. Records. — Trabut (1941), Jelenc (1955a): Kabylie de Collo, Dje-
bel Goufi. Feldmann (1947): Kabylie de Collo, Oued Elli Ankou,
Records. — Montagne (1838): Alger. Litschauer (1904), Jelenc 600 m. Jelenc (1955a): Kabylie du Djurdjura, Tala Ghizan. Bischler
(1954b, 1955a): Vallon bei la Bouzarea, 400 m. Trabut (1941), (2004): sine loco. Rare.
Bischler (2004): sine loco. Jelenc (1954b, 1967): Massif du Bou
Zegza, gorges de Keddara, 400 m. Jelenc (1955a): Kabylie des Ba-
bors, Cap Aokas; Kabylie du Djurdjura, Fort National; Monts de
Tlemcen, Forêt d’Hafir; Rouiba. Chevassut & Quézel (1956): Forêt Porella pinnata L.
de Pins d’Alep de Saint-Ferdinand. Chevassut & Quézel (1956),
Jelenc (1967): La Réghaïa. Jelenc (1956): Sahel d’Alger, au Beau- Records. — Lapie (1908), Jelenc (1955a): Kabylie du Djurdjura,
Fraisier. Jelenc (1967): Bainem. Frequent. Aizer. Very rare.

Porella platyphylla (L.) Pfeiff.

Phymatoceros bulbiculosus
(Brot.) Stotler, W.T.Doyle & Crand.-Stotl. Records. — Brongniart (1882): Blidah. Trabut (1888): sine loco.
Trabut (1941), Jelenc (1955a): Akfadou; Teniet-el-Haad. Jelenc
Records. — Trabut (1941), Jelenc (1955a): Aïn Taya; Alger. Che- (1949, 1951, 1955a, 1967): Monts de Tlemcem, Tlemcem. Trabut
vassut & Quézel (1956): Forêt de Pins d’Alep de Saint-Ferdinand; (1941), Jelenc (1951, 1955a): Monts de Tlemcen, Koudiat d’Hafir,
La Rassauta; La Réghaïa. Bischler (2004): sine loco. Rare. 1200 m. Jelenc (1955a): Kabylie du Djurdjura, Agoulmin Aberkane;
Kabylie du Djurdjura, Tala Ghizan. Rare.

Plagiochasma rupestre (J.R.Forst. & G.Forst.) Steph.

Radula complanata (L.) Dumort.
Records. — Trabut (1941), Jelenc (1955a): Bouçaada (Bou Saâda),
fissures des roches du Kerdada. Thériot (1933), Jelenc (1955a), Bis- Records. — Brongniart (1882), Corbière (1889), Jelenc (1955a):
chler (1978): Hoggar, au-dessus de l’Oued Ilaman, 2000-2100 m. Blida. Corbière (1904), Jelenc (1955a): Gorges de la Chiffa, ruisseau
Collenot et al. (1960): Tassili N’Aijer, Afaïtalèsse. Bischler (1978): des Singes. Trabut (1941): sine loco. Feldmann (1947): Kabylie de
Atlas saharien; Grand Kabylie; Petite Kabylie; Tassili, Plateau d‘Edjerit Collo, Oued Elli Ankou, 600 m. Jelenc (1955a): Djebel Chenoua;
E Djanet. Frahm (1978): Hoggar, Guelta von Imeghra, 1780 m. Kabylie de Collo, Djebel Goufi; Kabylie du Djudjura, Fôret d’
Bischler (2004): sine loco. Rare. Akfadou; Khodjaberry. Rare.

*Plagiochila asplenioides (L. emend. Taylor) Dumort. Radula lindenbergiana Gottsche ex C.Hartm.

Records. — Trabut (1941), Bischler (2004): sine loco. Jelenc (1951,

Comment 1955a, 1967): Monts de Tlemcen, Forêt de Zariffet, près de l’Aïn
Doubtfully present in Algeria. Due to the taxonomical prob- Defla, 1100 m. Very rare.
lems of this species (Söderström et al. 2002; Ros et al. 2007a),
probably the reports of this species in Kabylie du Djurdjura,
Afkadou by Trabut (1914, 1941) and Jelenc (1955a) belong Reboulia hemisphaerica (L.) Raddi
to P. porelloides. Records. — Brongniart (1882): Blidah. Corbière (1889): Blida,
glacière Laval. Corbière (1904), Jelenc (1955a): Kabylie, Fort Na-
tional, 1000 m; Souk-el-Haad. Corbière (1905), Jelenc (1955a):
Plagiochila porelloides Lindenb. (Torrey ex Nees) Environs de Constantine, les Pins; Environs de Constantine, Sidi
M’Cid. Lapie (1909): Kabylie du Djurjura. Maire & Sevenet (1928),
Jelenc (1955a): Gorges du Zaccar. Trabut (1941), Bischler (2004):
Records. — Jelenc (1955a): Kabylie du Djurdjura, Tala Ghizan. sine loco. Feldmann (1947), Jelenc (1955a): Kabylie de Collo, Djebel
Very rare. Goufi, 1000 m; Kabylie de Collo, vallée de l‘Oued Bin el Ouidante,
près de Taoulel, 600 m. Jelenc (1953): Atlas de Blida, Ravin Bleu de
Chréa; Bélezma, Tenient el Gantos, 1900 m. Jelenc (1953, 1955a,
Porella arboris-vitae (With.) Grolle 1967): Djurdjura, Plateau de l‘Aizer, 1900 m. Jelenc (1954b, 1967):
Chaîne des Zahrez, au nord du Zahrez Chergui, berges de l‘oued
Records. — Jelenc (1955a): Kabylie du Djudjura, Tala Ghizan; M‘Kamza au djebel Touillila, 1000 m. Jelenc (1955a): Ait Abdelali;
Kabylie du Djudjura, Yakouren. Rare. Ait Ouaban; Atlas de Blida, Oued el Kebir; Aurès, Djebel Ich Mah-

CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5) 59

 Ros R. M. et al.

mel, 2150 m; Djebel Chenoua; Edough; El Kantara; Forêt d‘Hafir; Riccia bicarinata Lindb.
Khenchela; Monts de Tlemcen, Tlemcen; Mouzaia; Ouarsenis, Téniet
el Had; Tirourda. Jelenc (1956): Atlas mitidjien, à Meurdja; Monts Records. — Trabut (1941), Jelenc (1955a): Alger à Aïn-Taya; Col
de Tlemcen, Forêt de Zariffet; Sahel d‘Alger, dans le ravin du Frais- de Sfa près Biskra; Oran. Chevassut & Quézel (1956): Forêt de Pins
Vallon. Jelenc (1959): Kabylie de Collo, Baie de Tamamart, 50 m. d’Alep de Saint-Ferdinand; La Rassauta; La Réghaïa. Jelenc (1967):
Jelenc (1966, 1967): Région de Boghar, sur le flanc nord du Djebel Kabylie du Djurdjura, Col de Tirourda. Jovet-Ast (1986), Bischler
Krellala, 1300 m. Jelenc (1967): Atlas de Blida; Massif du Bélezma, (2004): sine loco. Ros et al. (1990): Oued Rhiou. Rare.
Téniet el Gantès, 1900 m; Monts de Tlemcen; Yakouren. Ahayoun
et al. (2007): Djebel Ech chabir, vers Bou Arfa, 1400 m. Common.
Riccia bifurca Hoffm.
Riccardia chamedryfolia (With.) Grolle
Records. — Corbière (1904), Trabut (1941), Jelenc (1955a): La
Records. — Feldmann (1947), Jelenc (1955a): Kabylie de Collo, Bouzaréah, près Alger, 200 m. Corbière (1905): Environs de Con-
vallée de l’Oued Bin el Ouidante, près de Taoulel, 600 m; Kabylie stantine, Djebel Ouach, 900 m. Chevassut & Quézel (1956): La Ras-
de Collo, vallée de l’Oued Ramèche, 600-800 m. Trabut (1941), sauta; La Réghaïa. Jovet-Ast (1986), Bischler (2004): sine loco. Rare.
Jelenc (1955a): La Calle. Jelenc (1959): Kabylie de Collo, Baie de
Tamamart, 50 m. Rare.
Riccia canaliculata Hoffm.

*Riccardia latifrons (Lindb.) Lindb. Records. — Corbière (1904), Jelenc (1955a): Le Corso. Jovet-Ast
(1986): sine loco. Very rare.
Comment
Doubtfully present in Algeria. Bischler (2004) informed that Riccia cavernosa Hoffm.
it had been recorded previously in Algeria, but no published
reports have been found. Records. — Trabut (1941), Jovet-Ast (1986): Sahara, à In-Salah
(Massifs sahariens). Very rare.

Riccardia multifida (L.) Gray Comment

It was not listed for Algeria by Ros et al. (1999, 2007a), but at
Records. — Trabut (1914, 1941), Jelenc (1955a): Ruisseaux de la least one report was made by Trabut (1941) as Riccia crystal-
Forêt d’Akfadou. Trabut (1941): Kabylie. Jelenc (1955a): Kabylie du lina var. angustior Lindenb. Also, Bischler (2004) informed
Djurdjura, Yakouren. Jelenc (1956, 1966, 1967): Kabylie du Djurdjura,
Massif de l’Akfadou à la Fontaine des Ifs. Bischler (2004): sine loco. Rare. to it had been reported previously in Algeria. Rare.

Riccia atromarginata Levier Riccia chudoana Steph.

Records. — Trabut (1941): Mechouneche dans les Ziban. Jelenc Riccia chinensis Steph. ex Har. nom. nud.
(1955a): Aurès, M’chounech; Oran. Chevassut & Quézel (1956):
Forêt de Pins d’Alep de Saint-Ferdinand; La Réghaïa. Jelenc (1951, Records. — Hariot (1913), Jelenc (1955a): Tahount Arak (Mouy-
1967): Haute-Plaine de Tlemcen, collines de Bréa. Jovet-Ast (1986), dir). Stephani (1917), Jelenc (1955a): Sahara.
Bischler (2004): sine loco. Rare.
Comment
This species was not mentioned in the monograph of the genus
Riccia battandieri Trab. nom. nud. Riccia L. in the Mediterranean area by Jovet-Ast (1986), and
no other record or reference to it has been found (Ros et al.
Records. — Trabut (1941), Jelenc (1955a): Aïn-Taya (Alger). 2007a). Söderström et al. (2016) considered that there are
Chevassut & Quézel (1956): Forêt de Pin d’Alep de Bäinem; Forêt serious doubts about the value of the taxon. It was included
de Pin d’Alep d’El Riath. in Hodgetts et al. (2020) as a species occurring in territories
adjacent to Europe.
Comment
This name was proposed by Trabut (1941) from Algeria.
Grolle (1976) indicated that according to Jovet-Ast in litt., Riccia ciliata Hoffm.
it is a nomen dubium (Ros et al. 2007a). Riccia ciliata var. intumescens Bisch.
Riccia crinita auct. eur.
Riccia beyrichiana Hampe ex Lehm. Riccia intumescens (Bisch.) Underw.
Records. — Corbière (1904), Trabut (1941), Jelenc (1955a): La Riccia trichocarpa M.Howe
Bouzaréah, près Alger; Orléansville. Chevassut & Quézel (1956):
La Réghaïa. Jovet-Ast (1986): sine loco. Rare. Nomenclature follows Hodgetts et al. (2020).

60 CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5)

 Revisiting the Bryophyte Checklist of Algeria

Records. — Stephani (1885), Jovet-Ast (1986), Bischler (2004): and confirmed by Jovet-Ast (1957) as Riccia convexa is sit-
sine loco. Trabut (1941), Jelenc (1955a): Beni Khalfoum; Bône; uated in Mauritania (State Adrar, Region Aoujeft, County
Oran; Tiaret; Tlemcen. Trabut (1941), Jelenc (1955a), Chevassut & N’Teirguent, 19.71682N 12.86083W) and not to Algeria
Quézel (1956): Réghaia. Jelenc (1955a): Ouarsenis, Téniet el Had;
Rouiba; Saint Eugène; Tiguermount. Chevassut & Quézel (1956): as was stated by Ros et al. (2007a).
Forêt de Pins d’Alep de Saint-Ferdinand. Frequent.

Riccia glauca L. var. glauca

Riccia ciliifera Link ex Lindenb.
Records. — Montagne (1838), Trabut (1941), Jelenc (1955a): Alger.
Records. — Trabut (1941), Jelenc (1955a): Aïn-Taya, Bois sacré; Gandoger (1883), Jelenc (1967), Jovet-Ast (1986): sine loco. Cor-
Alger; Oran; Tiaret; Tlemcen. Jelenc (1955a), Chevassut & Qué- bière (1889), Jelenc (1955a): Blida, Chênes-Lièges. Corbière (1889):
zel (1956): La Réghaia. Chevassut & Quézel (1956): Forêt de Pins Corbière (1904), Jelenc (1955a): La Bouzaréah, près Alger. Jelenc
d’Alep de Saint-Ferdinand. Chevassut (1956): Atlas mitidjien, même (1956, 1967): Kabylie de Collo; Sersou, environs de Tiaret. Rare.
station que le 367. Müller (1957), Jovet-Ast (1986): sine loco. Jelenc
(1967): Barrage du Hamiz. Rare.
Riccia glauca var. ciliaris Warnst.

Riccia crozalsii Levier Records. — Trabut (1941): L’Agha. Trabut (1941), Jelenc (1955a):
Alger; Bouzaréah. Jelenc (1949, 1967): Haute-Plaine de Tlemcen,
Records. — Trabut (1941), Jelenc (1955a): Oran; Tlemcen à Hafir. colline de Bréa, 600 m. Rare.
Trabut (1941), Jelenc (1955a), Chevassut & Quézel (1956): Alger
à la Réghaia. Chevassut & Quézel (1956): La Rassauta. Jovet-Ast
(1986), Bischler (2004): sine loco. Rare. Riccia gougetiana Durieu & Mont. var. gougetiana

Records. — Durieu de Maisonneuve (1849), Trabut (1941), Jovet-Ast

Riccia crustata Trab. (1986), Bischler (2004): sine loco. Montagne (1856): Blida. Corbière
(1904), Jelenc (1955a): La Bouzaréah, près Alger. Corbière (1905),
Records. — Trabut (1916, 1941), Jelenc (1955a): Tiaret; Tlemcen Jelenc (1955a): Environs de Constantine, Djebel Ouach. Trabut
à Terni. Jovet-Ast (1986), Bischler (2004): sine loco. Rare. (1914), Jelenc (1955a): Forêt d’Akfadou. Müller (1941), Trabut
(1941), Jelenc (1955a): Alger. Trabut (1941), Jelenc (1955a): Tlemcen.
Jelenc (1951): Monts de Tlemcen, forêt de Zariffet, 1200 m. Jelenc
Riccia crystallina L. (1955a): Atlas de Blida; Chenoua. Jelenc (1955a), Chevassut & Qué-
zel (1956): La Réghaia. Chevassut & Quézel (1956): Forêt de Pins
Records. — Corbière (1904), Jelenc (1955a): Mostaganem. Litschau- d’Alep de Saint-Ferdinand. Ros et al. (1990): Oued Rhiou. Frequent.
er (1904), Jelenc (1955a): Fort National, 1000 m. Thériot (1933),
Jelenc (1955a): Hoggar, Ideles, 1500 m; Hoggar, Oued Ilaman,
2000 m; Hoggar, Tin Seghin, 2400 m, Mouydir, gorges d’Arak, 600- Riccia gougetiana var. armatissima
650 m; Tafedest, Oued Ahetes, 1150 m; Tassili N’Aijer, Amguid, Levier ex Müll. Frib.
750-800 m. Jelenc (1955a): Alger; Boufarik; Maison-Carrée; Monts
de Tlemcen, Forêt d’Hafir. Collenot et al. (1960): Tassili N’Aijer, Records. — Trabut (1914), Jelenc (1955a): Fort-National; Michelet.
Oued Ahalef. Jovet-Ast (1986), Bischler (2004): sine loco. Frequent. Trabut (1941), Jelenc (1955a): Berrouaghia, Montagne des Lions.
Trabut (1941): Oran. Trabut (1941), Jelenc (1955a): Tiaret. Trabut
(1941), Jelenc (1948, 1955a): Tlemcen. Jelenc (1948): Hafir. Jelenc
Riccia fluitans L. (1967): Monts de Tlemcen, 1200 m. Jovet-Ast (1986), Bischler
(2004): sine loco. Rare.
Records. — Trabut (1941), Jelenc (1955a): Fort de l’Eau. Trabut
(1941), Jelenc (1955a), Chevassut & Quézel (1956): Réghaia. Trabut
(1941): La Merdja. Jelenc (1955a): Inkermann, Oued Merja; Koléa;
Marais de la Rassauta; Marais de Senhadja. Jelenc (1967): Cheikh- Riccia huebeneriana Lindenb.
el-Bir. Jovet-Ast (1986): sine loco. Rare.
Records. — Maire (1939), Jelenc (1955a): Lac Freitis, dans la
plaine des Senhadja. Very rare.
Riccia frostii Austin
Riccia convexa Steph. Comment
Jovet-Ast (1986) and Bischler (2004) did not mention this species
Riccia convexa Steph. ex Har. nom. nud. to have been reported from Algeria, which was inded reported
Nomenclature follows Jovet-Ast (1957). under its synonym Riccia pseudo-frostii (Schiffn.) Müll. Frib.

Records. — Jovet-Ast (1986): Hoggar, Tassili des Ajjers. Very rare.

Riccia lamellosa Raddi
Comment
The locality Dhayet El-Tofla (Dayet el Tefla,), in the Sahara Records. — Corbière (1904): Mostaganem. Corbière (1904),
occidental, reported by Hariot (1913) and Jelenc (1955a), Trabut (1941): Orléansville, berges du Chélif et bois de la Pépin-

CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5) 61

 Ros R. M. et al.

ière. Trabut (1941): Oran. Trabut (1941), Jelenc (1955a): Alger. Riccia papillosa Moris
Jelenc (1949): Haute plaine de Tlemcen: colline de Bréa, 600 m.
Jelenc (1949, 1967): Monts de Tlemcen, cirque d’El Ourit, Records. — Corbière (1905), Trabut (1941), Jelenc (1955a): Envi-
700 m. Jelenc (1955a): Chercheell; Kabylie du Djurdjura, Adjiba; rons de Constantine, Djebel Ouach. Trabut (1941), Jelenc (1955a):
Oran, Massif du Murdjajo. Chevassut & Quézel (1956): Forêt Alger à la Réghaia. Jovet-Ast (1986), Bischler (2004): sine loco. Rare.
de Pin d’Alep de Bäinem; Forêt de Pin d’Alep d’El Riath; Forêt
de Pins d’Alep de Saint-Ferdinand. Jovet-Ast (1986): Massifs
sahariens. Jovet-Ast (1986), Bischler (2004): sine loco. Ros & Riccia perennis Steph.
Guerra (1987a): El Kantara. Ros et al. (1990): Oued Rhiou;
Ouled Mimoum. Frequent. Records. — Trabut (1916), Douin & Trabut (1919), Trabut (1941),
Jelenc (1955a), Chevassut & Quézel (1956): Fôret de la Réghaia
aux environs d’Alger. Chevassut & Quézel (1956): La Rassauta.
Riccia ligula Steph. Jovet-Ast (1986), Bischler (2004): sine loco. Rare.
Records. — Bischler (2004): sine loco. Very rare.
Riccia polycarpa (Trab.) Jelenc

Riccia macrocarpa Levier Records. — Trabut (1933), Jelenc (1955a): Hoggar, Asekrem,
gorges de Tin Segghin, 2400 m.
Records. — Trabut (1941): Tiaret. Trabut (1941), Jelenc (1955a):
Alger. Jelenc (1955a): Aïn Taya; La Bouzaréa; La Réghaia. Jovet-Ast
(1986), Bischler (2004): sine loco. Ros et al. (1990): Azazga. Rare. Comment
This species was not included in the monograph of the genus
Riccia L. in the Mediterranean area by Jovet-Ast (1986), but
Riccia mamillata Trab. ex Steph. was listed by Söderström et al. (2002, 2016), who mentioned
knowledge problems. It was included in Hodgetts et al. (2020)
Records. — Stephani (1889b): sine loco. Trabut (1941), Jelenc as a species occurring in territories adjacent to Europe.
(1955a): El-Adjiba (Alger).

Comment Riccia sommieri Levier

This species was not included in the monograph of the genus
Riccia L. in the Mediterranean area by Jovet-Ast (1986), but Records. — Trabut (1941), Jelenc (1955a): Tiaret. Chevassut &
Quézel (1956): Forêt de Pin d’Alep de Bäinem; Forêt de Pin d’Alep
was listed by Söderström et al. (2002, 2016), who mentioned d’El Riath. Jovet-Ast (1986), Bischler (2004): sine loco. Rare.
knowledge problems. It was included in Hodgetts et al. (2020)
as a species occurring in territories adjacent to Europe. Accord-
ing to Casares-Gil (1919) it is similar to Riccia macrocarpa Riccia sorocarpa Bisch.
Levier. Its identity remains doubtful.
Riccia sorocarpa var. heigii Schiffn.
Nomenclature follows Hodgetts et al. (2020).
Riccia michelii Raddi
Records. — Corbière (1904), Trabut (1941), Jelenc (1955a): Alger.
Records. — Corbière (1904), Jelenc (1955a): Kabylie, Fort Na- Corbière (1904), Jelenc (1955a): Kabylie, Fort National, 1000 m;
tional, 1000 m; Mostaganem, 170 m; Oran. Corbière (1905), Mostaganem; Oran. Corbière (1905), Jelenc (1949): Colline de Bréa.
Jelenc (1955a): Environs de Constantine, Djebel Ouach. Trabut Jelenc (1955a): Environs de Constantine, Djebel Ouach. Trabut (1941),
(1941), Jovet-Ast (1986), Bischler (2004): sine loco. Jelenc (1951): Jelenc (1955a): Kabylie du Djurdjura, Boghni. Jelenc (1955a): Aïn Taya;
Haute plaine de Tlemcen, collines de Bréa. Jelenc (1955a): Aïn Rouiba. Jelenc (1955a), Chevassut & Quézel (1956): La Réghaia. Jelenc
Taya; Kabylie du Djurdjura, Akfadou; Ouarsenis, Téniet el Had. (1951, 1967): Haute-Plaine de Tlemcen, 600 m. Jovet-Ast (1986):
Jelenc (1955a, 1956), Chevassut & Quézel (1956): La Réghaia. Massifs sahariens. Jovet-Ast (1986), Bischler (2004): sine loco. Frequent.
Jelenc (1955a), Ahayoun et al. (2007): La Bouzaréa. Jelenc (1956,
1967): Atlas mitidjien, au barrage du Hamiz. Chevassut & Quézel
(1956): Forêt de Pin d’Alep de Bäinem; Forêt de Pin d’Alep d’El Riccia subbifurca Warnst. ex Croz.
Riath; Forêt de Pins d’Alep de Saint-Ferdinand; La Rassauta. Jelenc
(1967): Haute-Plaine de Tlemcen, 600 m. Frequent. Records. — Trabut (1941): Alger. Trabut (1941): Jelenc (1955a):
La Réghaia. Jovet-Ast (1986), Bischler (2004): sine loco. Rare.

Riccia nigrella DC. Riccia trabutiana Steph.

Records. — Corbière (1904), Jelenc (1955a): La Bouzaréah, Records. — Stephani (1889b), Jovet-Ast (1986), Bischler (2004):
près Alger, 200 m; Mostaganem, 150 m. Corbière (1905), Jelenc sine loco. Trabut (1941): Blida; Montagnes du sud Oranais. Jelenc
(1955a): Environs de Constantine, Djebel Ouach. Trabut (1941), (1949, 1951, 1967): Haute plaine de Tlemcen, collines de Bréa.
Jelenc (1955a): Alger; Tiaret. Jelenc (1955a): Aïn Taya; Cap Matifou; Jovet-Ast (1955), Jelenc (1955a): Oued el Kebir, à Blida. Jelenc
La Réghaia. Jovet-Ast (1986), Bischler (2004): sine loco. Ros et al. (1955a): Nemours. Chevassut & Quézel (1956): Forêt de Pins
(1990): Azazga; Dellys, 0-40 m; Oued Rhiou. Frequent. d’Alep de Saint-Ferdinand. Rare.

62 CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5)

 Revisiting the Bryophyte Checklist of Algeria

Riccia warnstorfii Limpr. ex Warnst. Riella macrocarpa (P.Allorge)

Puche, Segarra-Moragues, Sabovlj., M.Infante & Heras
Riccia warnstorfii var. subinermis Warnst.
Records. — Segarra-Moragues et al. (2022): sine loco. Very rare.
Nomenclature follows Hodgetts et al. (2020).
Records. — Trabut (1941), Jelenc (1955a): Alger. Jelenc (1951), Comment
Jelenc (1967): Monts de Tlemcen, forêt de Zariffet, vers 1150 m,
l’oued Zarifète. Jelenc (1955a): Aïn Taya. Jovet-Ast (1986), Bischler The species was recently recognized by Segarra-Moragues et al.
(2004): sine loco. Rare. (2022) for the previously considered Riella helicophylla var.
macrocarpa P.Allorge. They reported just an Algerian sterile
clone obtained from U. Kutschera, the species is therefore to
Ricciocarpos natans (L.) Corda be found wild in the country. According to these authors, it
extends from the western Mediterranean in Portugal to the
Records. — Trabut (1941): Alger; Entre Blida et Coléa; Fort-de-
l’Eau. Trabut (1941), Jelenc (1955a): Fossés de Farghen; Marais de
Eastern Mediterranean in Cyprus, replacing in many, but not
la Rassauta. Rare. all, countries the reports of R. helicophylla.

Riella affinis M.Howe & Underw. Riella notarisii (Mont.) Mont.

Records. — Puche & Boisset (2009), Segarra-Moragues et al. Riella battandieri Trab.
(2014): Tassili N’Ajjer, Tafilalet canyon, 24°36’05”N, 9°37’26”E,
1600 m. Very rare. Riella reuteri Mont.
Riella sersuensis Trab.
Riella bialata Trab.
Nomenclature follows Segarra-Moragues et al. (2022).
Records. — Trabut (1908, 1941), Jelenc (1955a): Plaine du Chéliff, Records. — Trabut (1886, 1941), Jelenc (1955a, 1957): In fossis
Duperré. Allorge (1932): sine loco. Very rare. hieme inundatis circa vicum Maison-Blanche Alger (Plaine de la
Mitidja, Alger à Maison Carrée, fossé de l’Oued Smar). Allor-
Riella cossoniana Trab. ge (1932): sine loco. Trabut (1934, 1941), Jelenc (1955a, 1956,
1957, 1967): Sersou, autour de la mare de Tasselent (entre Tiaret
et Vialar). Jelenc (1957): Vallé de l’oued Chélif (Ouest Algerian).
Records. — Battandier & Trabut (1886), Trabut (1887b, 1941), Jelenc Chevassut & Quézel (1958), Jelenc (1967): Entre la ferme Bourlier
(1955a, 1957): Chott Chergui, El Kreider. Allorge (1932): sine loco. et Tiaret, à 20 km à l’E de Tiaret, de part de d’autre de la route
Maire (1937), Jelenc (1955a): Sahara central, Hoggar, Tamanghasset. Tiaret-Vialar. Rare.
Jelenc (1957): Sud-Oranais, Khunda. Frahm (1978): El Goléa. Rare.

Comment
Comment
According to Jelenc (1957), the locality “Sud-Oranais, Khunda”
Riella battandieri and R. reuteri were considered by Jelenc
appears in the label of the sample collected by Trabut in
(1957) synonyms of R. notarisii (Mont.) Mont. That was
April 1907, at PC; he indicates that he did not locate any
also the criterion of Schumacker et al. (2005) and followed
geographical place with this name and that this word is very
by Ros et al. (2007a). Although both species were listed
similar to a word in the Arabic language “Khandek”, and that
by Söderström et al. (2016) and Hodgetts et al. (2020) as
it was possibly a mistake to refer to the locality “Le Kreider”.
accepted species, these works also stated that their taxo-
nomical distinction is questioned and that they belong to
Riella notarisii complex with poorly defined taxonomic
Riella helicophylla (Bory & Mont.) Mont.
boundaries, as did Segarra-Maragues et al. (2022). Jelenc
Records. — Bory de Saint-Vincent & Montagne (1843), Mon- (1957) stated that the area where R. notarisii had been found
tagne (1856), Trabut (1941), Jelenc (1955a), Segarra-Moragues in Algeria (sub R. battandieri) had been cultivated at that
et al. (2022): près d’un petit lac d’eau saumâtre (La Sénia) situé à time and it should be supposedly taken that the species
environ 8 km au S-E d’Oran. Durieu de Maisonneuve (1846): sine had disappeared. The only reference to R. reuteri in Algeria
loco. Cosson (1853): D’Oran au Chott-el-Chergui. Trabut (1887a):
Chott (Khreider). Jelenc (1957): Daya Borselly, Oran, Munby. Jelenc was done by Allorge (1932), in a map of his Pflanzenareale
(1957), Segarra-Moragues et al. (2022): Oran dans le Lac Dahia el series, but no previous report seems to have been published
Morceli. Jelenc (1967): Mécheria; Miserghin; Valmy. Segarra-Mor- on this name.
agues et al. (2022): Hochland der Chotts, Süsswassersee südl. Ksar
El Boukhari bei Bougzoul. Rare.

Comment Riella numidica Trab.

The study of Segarra-Moragues et al. (2022) confirmed that the Records. — Trabut (1934, 1941), Jelenc (1955a, 1957): Source à
species is restricted to the western Mediterranean basin, being Boniou (Chott Hodna). Allorge (1932): sine loco. Jelenc (1955a):
present in Algeria, Morocco, Tunisia, and mainland Spain. Oran, Sainte Barbe du Tlélat. Rare.

CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5) 63

 Ros R. M. et al.

Riella parisii Gottsche Solenostoma gracillimum (Sm.) R.M.Schust.

Records. — Durieu de Maisonneuve (1868), Jelenc (1957), Se- Jungermannia gracillima Sm.
garra-Moragues et al. (2022): Fossés d’obstacle près de Berbessa
(Plaine de la Mitidja, Bassin de l’Oued Mazafran, Barbessa, à la Nomenclature follows Söderström et al. (2016).
limite du Sahel et de la Mitidja, 2°45’E, 36°37’N, 30 m). Bat-
Records. — Corbière (1904): La Calle, route de Tabarka, entre
tandier & Trabut (1886), Trabut (1941), Jelenc (1955a, 1957):
Oum Theboul et El Aioun, 400 m. Trabut (1941), Jelenc (1955a):
Plaine de la Mitidja, Boufarick (2°55’E, 36°35’N); Plaine de la
La Calle. Feldmann (1947), Jelenc (1955a): Forêt de Taourirt
Mitidja, Bassin de l’oued Mazafran, Chaïba inférieur et Farghen
Ighil, Grande Kabylie; Kabylie de Collo, Oued Elli Ankou, 500-
(2°50’E, 36°37’N, 13 m); Plaine de la Mitidja, Bassin de l’Oued
900 m. Feldmann (1947), Jelenc (1955a), Váňa (1974): Kabylie
el Harrach, environs de Maison-Carrée, Alger (3°8’E, 36°43’N,
de Collo, vallée de l’Oued Ramèche, 500-900 m. Bischler (2004):
15 m). Allorge (1932): sine loco. Jelenc (1955a, 1957), Chevassut
sine loco. Rare.
(1956): Plaine de la Mitidja, Bassin de l’Oued el Hamiz, marais
de la Rassauta (3°12’E, 36°45’N, 5 m). Jelenc (1955a): Sebkhra
d’Oran. Jelenc (1955a, 1957): Plaine de la Mitidja, bassin de
l’Oued el Réghaïa, nord de la ville de Réghaïa (3°20’E, 36°47’N,
20 m); Sahel de Koléa, environs de Coleah, Bou Ismail, massif du
Solenostoma hyalinum (Lyell) Mitt.
Chenoua (2°45’E, 36°40’N). Jelenc (1957): Plaine de la Mitidja,
Bassin de l’oued Mazafran, Chaïba inférieur et Oued el Alleug, Jungermannia hyalina Lyell
2°45’E, 36°35’N, 30 m. Jelenc (1957), Segarra-Moragues et al. Nomenclature follows Söderström et al. (2016).
(2022): Plaine de la Mitidja, environs d’Alger. Jelenc (1957),
Ahayoun et al. (2007): Plaine de la Mitidja, Oued el Alleg, 2°48’E, Records. — Corbière (1904): Alger, chemin de Telemly, 80 m.
36°33’N. Rare. Trabut (1941), Jelenc (1955a): Alger. Trabut (1914, 1941): Kabylie.
Jelenc (1955a): Atlas de Blida. Rare.

Scapania compacta (A.Roth) Dumort.

*Solenostoma obovatum (Nees) C.Massal.
Records. — Trabut (1941), Jelenc (1955a): La Calle. Feldmann
(1947), Jelenc (1955a): Kabylie de Collo, Taoulel. Bischler (2004): Jungermannia obovata Nees
sine loco. Rare.
Nomenclature follows Söderström et al. 2016.

Scapania curta (Mart.) Dumort. Comment

Doubtfully present in Algeria. Bischler (2004) informed that
Records. — Trabut (1941), Jelenc (1955a): Akfadou. Very rare. it had been recorded previously in Algeria, but no published
reports have been found.

*Scapania gracilis Lindb. Solenostoma sphaerocarpum (Hook.) Steph.

Comment Jungermannia sphaerocarpa Hook.
Doubtfully present in Algeria. Bischler (2004) informed that Nomenclature follows Söderström et al. (2016).
it had been recorded previously in Algeria, but no published
reports have been found. Records. — Feldmann (1947), Jelenc (1955a): Kabylie de Collo,
Maison forestière de Bou Noghra, 600 m. Very rare.

Scapania irrigua (Nees) Nees Southbya nigrella (De Not.) Henriq.

Records. — Jelenc (1955a): Kabylie du Djurdjura, Yakouren. Records. — Corbière (1904), Jelenc (1955a): Orléansville, bois
Very rare. de la Pépinière. Corbière (1905), Jelenc (1955a): Environs de
Constantine, Mansourah. Trabut (1941), Bischler (2004): sine loco.
Jelenc (1949, 1967): Monts de Tlemcen, flanc nord du djebel el
Scapania subalpina (Nees ex Lindenb.) Dumort. Béniane, 1000 m. Rare.

Records. — Maire (1940), Jelenc (1955a): Kabylie, ruisselets dans

les chênaies de l’Akfadou, sur les grés, 1300-1400 m. Very rare. Southbya tophacea (Spruce) Spruce
Records. — Corbière (1904), Jelenc (1955a, 1967): Alger, 80 m.
Scapania undulata (L.) Dumort. Corbière (1905), Jelenc (1955a): Environs de Constantine, Chet-
tabah. Corbière (1905): Environs de Constantine, les Pins. Trabut
Records. — Trabut (1914): Djurdjura. Trabut (1941), Jelenc (1941), Bischler (2004): sine loco. Jelenc (1954b): Sahel d‘Alger,
(1955a): Akfadou; Kabylie. Jelenc (1955a): Kabylie du Djurdjura, colline du Hamma. Jelenc (1955a): Arba; Atlas de Blida, 1200 m;
Ait Idjer. Jelenc (1967): Kabylie du Djurdjura, Yakouren. Bischler La Bouzaréa; Marengo. Jelenc (1956): Sahel d’Alger, dans le ravin
(2004): sine loco. Rare. du Frais-Vallon. Ros et al. (1990): Chiffa’s Gorge. Rare.

64 CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5)

 Revisiting the Bryophyte Checklist of Algeria

Sphaerocarpos michelii Bellardi Environs de Constantine, djebel Ouach. Jelenc (1955a): Alma;
Maison-Carrée. Ros et al. (1990): Dellys, 0-40 m. Boukhatem
Records. — Corbière (1904), Jelenc (1955a): Mostaganem, 104 m. et al. (2017): El-Kala National Park, Tonga lake watershed. Rare.
Trabut (1941), Bischler (2004): sine loco. Jelenc (1954b, 1956):
Sahel d‘Alger, colline du Hamma. Jelenc (1955a, 1967): Alger. Je-
lenc (1955a): Kabylie du Djurdjura, Akfadou; Philippeville. Rare.
Acaulon triquetrum (Spruce) Müll. Hal.

Sphaerocarpos texanus Austin Records. — Bescherelle (1882): Alger, au-dessus de Mustapha,

Bab-Azoun; Mustapha. Bescherelle (1882), Jelenc (1955a): Aïn-Va-
Records. — Trabut (1941): L‘Agha. Jelenc (1955a): Alger. Bischler gout; Alger. Corbière (1904): Mostaganem. Corbière (1904), Jelenc
(2004): sine loco. Rare. (1955a): Sétif, 1112 m. Corbière (1905), Jelenc (1955a): Environs
de Constantine, Chettabah; Environs de Constantine, djebel Ou-
ach. Ros et al. (1990): Ouled Mimoum. Boukhatem et al. (2017):
Comment El-Kala National Park, Tonga lake watershed. Rare.
Bell et al. (2013) showed that the European populations of
Sphaerocarpos texanus are genetically very distinct from the
American populations, and that S. europaeus Lorb. should be Alleniella complanata (Hedw.)
used for the European species. Studies in northern African S.Olsson, Enroth & D.Quandt
populations should be made to confirm if they also corre-
spond to S. europaeus. Records. — Bescherelle (1882): Djurjura. Bescherelle (1882),
Recently, Infante et al. (2024) described a new species in the Jelenc (1955a): Alger, forêt de Chaïba, à oued-el-Alleug; La Calle;
Philippeville. Trabut (1888): sine loco. Corbière (1905): Environs de
Iberian Peninsula (Sphaerocarpos ibericus M.Infante, Luceño, Constantine, oued Marsa. Jelenc (1955a): Alger; Boufarik; Kabylie
Quirós-de-la-Peña, Míguez, J.Muñoz & Heras), which could des Babor, Tababor; Kabylie du Djurdjura, Akfadou; Kabylie du
also be present in Algeria. Djurdjura, Izer Zouggaren; Khodjaberry. Ahayoun et al. (2007):
Kabylie. Frequent.

Targionia hypophylla L.
Aloina aloides (Koch ex Schultz) Kindb.
Records. — Montagne (1838), Jelenc (1955a), Ahayoun et al.
(2007): Alger. Brongniart (1882): Blidah. Gandoger (1883), Bischler Records. — Bescherelle (1882): Alger; Route de Saint-Eugène.
(2004): sine loco. Trabut (1887a): D’Oran a Mécheria. Corbière Bescherelle (1882), Corbière (1889): Blidah. Corbière (1904): Alger,
(1889): Blida, Chemin de la glacière Laval. Litschauer (1904), Jelenc talus à Telemly. Corbière (1904), Jelenc (1955a): Médéa. Corbière
(1955a): Fort National, 1000 m; Im Tal des Qued-Kebir bei Blidah, (1905): Environs de Constantine, route du djebel Ouach; Environs
200 m. Corbière (1905): Environs de Constantine. Lapie (1909): de Constantine, route du Moulin Laire. Jelenc (1949, 1967): Monts
Kabylie du Djurjura. Maire & Sevenet (1928), Jelenc (1955a): Alger, de Tlemcen, djebel el Béniane, 1100 m. Jelenc (1955a): Alger, Frais
Tombeau de la Chrétienne, près de Bérard. Jelenc (1949): Djebel Vallon, 300 m; Atlas de Blida, oued el Kebir, 200 m; Constantine;
Murdjadjo (Oran), Ravin du Tammermouth; Vallée de l’Isser “El Kabylie du Djurdjura, Aït Aïcha; Kabylie du Djurdjura, Aït Idjer;
Kasbat”, 500 m. Jelenc (1949, 1967): Monts et Haute-Plaine de Monts de Tlemcen, Mazer. Jelenc (1966, 1967): Atlas Mitidjien à
Tlemcen. Jelenc (1951, 1967): Massif du Tessala, pic de Tafaraoui, Hammam Melouane. Delgadillo (1975): Alger, vallée des Consults.
600 m. Jelenc (1955a): Birmandreis; Edough; El Biar; Gorges de la Ros & Guerra (1987a): El Kantara. Ros et al. (1990): Between Mo-
Chiffa; La Bouzaréa; Monts de Constantine; Monts de Tlemcem, staganem and Cap Ivi, Abdelmaimalek-Ramdan beach, 0 m; Oued
cascades d’El Ourit; Oran, Ravins de Noiseaux. Jelenc (1967): Rhiou; Ouled Mimoum. Gallego et al. (1999): Alger-Vablei, des
Kabylie de Collo; Kabylie du Djurdjura, Yakouren; Sahel d’Oran. eonseds. Common.
Ros & Guerra (1987a): El Kantara. Ros et al. (1990): Between
Mostaganem and Cap Ivi, Abdelmaimalek-Ramdan beach, 0 m;
Djebel Chenoua, 5-40 m; Oued Rhiou. Common.
Aloina ambigua (Bruch & Schimp.) Limpr.

Records. — Bescherelle (1882): Bône; Kabylie. Bescherelle (1882),

Targionia lorbeeriana Müll. Frib. Jelenc (1955a): Alger; Kouba; Maison-Carrée; Oran; Tlemcen.
Gandoger (1883): sine loco. Trabut (1887a): D’Oran à Mécheria.
Records. — Jelenc (1956, 1967): Massif de la Bouzaréa, sous les Corbière (1889): Blida. Corbière (1905): Environs de Constantine,
Pins dans la foret de Baïnem. Very rare. La Corniche; Environs de Constantine, Salah Bey. Lapie (1909):
Kabylie du Djurjura. Jelenc (1949): Plaine d’Oran, talus de fossé à
La Sénia. Jelenc (1951): Sahel d’Arzeu, djebel Borouse. Jelenc (1951,
Mosses 1967): Haute-Plaine et Monts de Tlemcen. Jelenc (1954b): Chaîne
des Zahrez, au nord du Zahrez Chergui, près de l’Aïn Hallouf au
djebel Touilila, 1000 m. Jelenc (1955a): Atlas de Blida, oued el Kebir;
Acaulon muticum (Hedw.) Müll. Hal. Birmandreïs; Frais Vallon; Gorges de la Chiffa; Monts de Constantine;
Ouarsenis, djebel Doui. Jelenc (1966, 1967): Massif du Tessala, pic
Records. — Bescherelle (1882): Birkadem; Pointe-Pescade. Be- Tessala, 1000 m. Jelenc (1966): Sahel d’Arzeu à Assi bou Nif. Jelenc
scherelle (1882), Jelenc (1955a, 1967): Alger. Bescherelle (1882), (1967): La Sénia, Assi-bou-Nif. Bizot (1973): Ruines de Timgad.
Jelenc (1955a, 1955b, 1967): La Calle. Corbière (1905): Environs Delgadillo (1975): Tizi-Ouzou. Gallego et al. (1999): Kabylie de
de Constantine, route de Batna. Corbière (1905), Jelenc (1955a): Collo de Constantine, estuarie de L’oued Guebli Collo. Common.

CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5) 65

 Ros R. M. et al.

Aloina brevirostris (Hook & Grev.) Kindb. Archidium alternifolium (Hedw.) Mitt.

Records. — Gallego et al. (1999): Oran. Very rare. Records. — Bescherelle (1882), Jelenc (1955a), Snider (1975):
La Calle, bruyères; Philippeville. Jelenc (1955a): Alger; La Bou-
zaréa. Boukhatem et al. (2017): El-Kala National Park, Tonga lake
watershed. Rare.
Aloina rigida (Hedw.) Limpr.

Records. — Montagne (1838), Jelenc (1955a): Alger. Bescherelle Aschisma carniolicum (F.Weber & D.Mohr) Lindb.
(1882), Jelenc (1955a): Cap Matifou; Constantine; Gigthi. Gallego
et al. (1999): Oran. Rare. Records. — Corbière (1905), Jelenc (1955a): Environs de Con-
stantine, djebel Ouach. Jelenc (1955a): Alger, La Réghaia. Rare.
Amblystegium serpens (Hedw.) Schimp.
Atrichum undulatum (Hedw.) P.Beauv.
Records. — Bescherelle (1882): sine loco. Feldmann (1947), Jelenc
(1955a): Kabylie de Collo, Bessombourg, 500-600 m. Very rare. Records. — Bescherelle (1882), Jelenc (1955a): Djebel Édough.
Jelenc (1955a): Alger; Kabylie du Djurdjura, Agoulmin Aberkan;
Kabylie du Djurdjura, forêt d’Akfadou. Rare.
Anacolia webbii (Mont.) Schimp.

Records. — Bescherelle (1882): Blidah, aux Deux-Frères; Kabylie; Aulacomnium androgynum (Hedw.) Schwägr.
La Mouzaïa. Litschauer (1904), Jelenc (1955a): Qued-Kebir bei
Blidah, 200 m. Trabut (1914): Djurdjura. Jelenc (1955a): Atlas Records. — Jelenc (1955a): Djurdjura; Monts de Constantine,
de Blida, 1400 m; Kabylie du Djurdjura, Akfadou; Kabylie du djebel Ouach; Ouarsenis, Téniet el Had. Rare.
Djurdjura, Azerou Arzela; Kabylie du Djurdjura, Tala Ghizan. Rare.

Aulacomnium palustre (Hedw.) Schwägr.

Anomobryum julaceum
(Schrad. ex P.Gaertn. et al.) Schimp. Records. — Bescherelle (1882), Trabut (1914), Jelenc (1955a):
Djurjura. Jelenc (1955a): Monts de Constantine; Ouarsenis, Téniet
Records. — Bescherelle (1882): Birkadem; Ravins, près le Fort el Had. Rare.
l’Empereur. Bescherelle (1882), Trabut (1914), Jelenc (1955a):
Col de Tirourda. Bescherelle (1882), Jelenc (1955a): Alger, talus
des chemins, au Bouzaréah. Jelenc (1955a): Constantine, 700 m; Barbula unguiculata Hedw.
Kabylie du Djurdjura, Akfadou; Kabylie du Djurdjura, Fort Na-
tional; Kabylie du Djurdjura, Michelet, 1000-1200 m; Kabylie du Records. — Montagne (1838), Bescherelle (1882), Jelenc (1955a):
Djurdjura, Tifrit; Philippeville. Rare. Alger. Bescherelle (1882), Jelenc (1955a): Birmandreïs; Bône; Édough;
Kouba; Tiaret. Gandoger (1883): sine loco. Corbière (1889): Blida.
Corbière (1904): Médéa. Corbière (1904), Jelenc (1955a): Gorges
de la Chiffa, ruisseau des Singes. Corbière (1905): Environs de
Anomodon viticulosus (Hedw.) Hook. & Taylor Constantine, Chettabah; Environs de Constantine, Menidj; En-
virons de Constantine, Pépinière du djebel Ouach; Environs de
Records. — Bescherelle (1882): Kabylie. Bescherelle (1882), Jelenc Constantine, près l’oued Bou Merzoug; Environs de Constantine,
(1955a): Djebel Tababor; Taguemount-Azouz. Rare. tuileries du Rummel. Braun-Blanquet & Maire (1924), Jelenc
(1955a): Tlemcen. Jelenc (1955a): Atlas de Blida, oued el Kebir,
200 m; Dellys; Kabylie du Djurdjura, Fort National; La Réghaia.
Antitrichia californica Sull. Ros et al. (1990): Between Mostaganem and Cap Ivi, Abdelmei-
malek-Ramdan beach, 0 m; Oued Rhiou; Ouled Mimoum. Cano
Records. — Bescherelle (1882): Alger, Atlas; Mont de l’Ouarsenis; et al. (2000): Kasba Cheik Saïd. Boukhatem et al. (2017): El-Kala
Serdj-el-Aouda. Bescherelle (1882), Jelenc (1955a): Aïn-Talazid; Forêt de National Park, Tonga lake watershed. Mazari & Boulaacheb (2023):
cèdres de Teniet-el-Haâd. Brongniart (1882): Blidah. Corbière (1904), Setif, Megriss Mountain. Common.
Jelenc (1955a): Djebel Mouzaïa. Litschauer (1904), Jelenc (1955a):
Von Fort National nach Michelet, 1000-1200 m. Jelenc (1951): Forêt
de Zariffet, 1100 m. Jelenc (1951, 1967): Monts de Tlemcen, Forêt de Bartramia aprica Müll. Hal.
Tessera Mramet, 1400 m. Jelenc (1955a): Kabylie du Djurdjura, Aït
Ouaban; Kabylie du Djurdjura, Beni Khalfoun; Kabylie du Djurdjura, Bartramia stricta auct. eur.
Tala Ghizan; Massif de la Mahouna, Serdj el Aouda; Ouarsenis, dje-
bel Doui. Jelenc (1967): Kabylie de Collo, dejbel Goufi. Boukhatem Bartramia rosamrosiae Damayanti, J.Muñoz, J.-P.Frahm & D.Quandt
et al. (2017): El-Kala National Park, Tonga lake watershed. Frequent.
Nomenclature follows Müller (2014).
Records. — Montagne (1838), Bescherelle (1882), Jelenc (1955a):
Antitrichia curtipendula (Hedw.) Brid. Alger. Bescherelle (1882): Couteaux, Bouzaréah; Djurjura; Kabylie;
La Galite; L’Agha. Bescherelle (1882), Corbière (1889): Blidah.
Records. — Jelenc (1955a) Djurdjura, Aït Ouaban; Djurdjura, Bescherelle (1882), Jelenc (1955a): Bône; La Calle; Mascara;
Fort National. Rare. Philippeville; Tiaret. Gandoger (1883): sine loco. Trabut (1887a):

66 CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5)

 Revisiting the Bryophyte Checklist of Algeria

D’Oran à Mécheria, Tell. Corbière (1904): Bône, rochers siliceux Brachymenium exile
au pied de l’Édough. Corbière (1904, 1905), Jelenc (1955a): (Dozy & Molk.) Bosch. & Sande Lac.
Constantine, bois du djebel Ouach. Litschauer (1904): Col de
Tirourda, 1700 m; Fort National und Michelet, 1000-1200 m.
Litschauer (1904), Jelenc (1955a): Chiffaschlucht und im Tal Records. — Thériot (1931, 1933), Jelenc (1955a): Atakor-n-
des Qued-Kebir bei Blidah, 300 m. Lapie (1909): Kabylie du Ahaggar, Ilaman, 2000 m. Thériot (1931, 1933), Jelenc (1955a),
Djurjura. Cardot (1904), Maire (1916): Province d’Oran, Djebel Ochi (1972): Ideles, 1500 m; Tefedest, 1200 m. Rare.
Aïssa, 1800 m, 2250 m. Jelenc (1949, 1967): Monts de Tlemcen,
forêt de Zariffet, 1150 m; Djebel Chenoua. Jelenc (1951, 1967):
Massif du Tessala, pic de Tafaraoui, 650 m; Sahel d’Oran, plateau Brachytheciastrum salicinum
des Ghamras, 400 m. Jelenc (1954b, 1967): Atlas mitidjien, fôret (Schimp.) J.D.Orgaz, M.J.Cano & J.Guerra
d’El Aioun, 500 m; Massif de Bou Zezga, gorges de Keddara,
400 m; Zahrez Chergui, djebel Touilila, 1000 m. Jelenc (1955a): Brachytheciastrum velutinum var. salicinum (Schimp.) Ochyra &
Akfadou; Col des Aït Aïcha; Djebel Doui; Kabylie des Babor, cap Żarnowiec
Aokas; Kabylie du Djurdjura, Aït Khalfoun; Kabylie du Djurdjura,
Bouzgeur; Kabylie du Djurdjura, col de Tirourda, 1700 m; Kabylie Nomenclature follows Orgaz et al. (2013).
du Djurdjura, Fort National, Michelet, 1000-1200 m; La Bouza-
réa; Monts des Ksour, djebel Aïssa, 1800 m; Ouarsenis; Tablabalt; Records. — Jelenc (1953): Atlas de Blida, ravin Bleu de Chréa;
Taourirt Aberkane. Jelenc (1955a), Ros et al. (1990): Gorges de la Aurès, forêt de Sgag, 1700 m. Jelenc (1953, 1955a, 1967): Ba-
Chiffa. Jelenc (1955a), Ahayoun et al. (2007): Djebel Chenoua. bor, flanc nord du Babor, 1700 m. Jelenc (1953, 1967): Bèlezma,
Jelenc (1956): Massif de la Bouzaréa, forêt de Baïnem; Massif du cédraies de l’oued Chaba, 1400 m. Jelenc (1955a): Atlas de Blida,
Tessala, pic Tessala. Jelenc (1967): Kabylie de Collo. Ros et al. forêt de Quercus; Kabylie des Babor, cap Aokas; Monts de Con-
(1990): Texenna’s wood, 700 m. Boukhatem et al. (2017): El-Kala stantine, djebel Ouach; Ouarsenis, Téniet el Had. Jelenc (1967):
National Park, Tonga lake watershed. Very common. Massif de l’Aurès, djebel Chélia, 1800 m; Massif de l’Aurès, djebel
Pharounm, 1800 m; Massif de l’Aurès, forêt de Sgag, 1700 m.
Frequent.
Bartramia pomiformis Hedw.
Records. — Desfontaines (1799): sine loco. Bescherelle (1882): Brachytheciastrum trachypodium
Kabylie. Bescherelle (1882), Jelenc (1955a): Blidah, Atlas. Corbière (Brid.) Ignatov & Huttunen
(1889): Blida, Tala-Yesid. Litschauer (1904), Jelenc (1955a): Col
de Tirourda, 1700 m. Feldmann (1947), Jelenc (1955a): Kabylie
Records. — Corbière (1889), Jelenc (1955a): Blida, Koudiat-Chréa,
de Collo, sommet du djebel Goufi, versant Nord, 1000 m. Jelenc
1500 m. Corbière (1904), Jelenc (1955a): Médéa. Rare.
(1955a): Bône; Frais Vallon; Gorges de la Chiffa; Kabylie du Djurd-
jura, Bouzgeur. Jelenc (1955a, 1956, 1967): Kabylie du Djurdjura,
Akfadou. Boukhatem et al. (2017): El-Kala National Park, Tonga
lake watershed. Frequent. Brachytheciastrum velutinum
(Hedw.) Ignatov & Huttunen

*Brachymenium commutatum (Müll. Hal.) A.Jaeger Records. — Bescherelle (1882): Kabylie. Bescherelle (1882),
Jelenc (1955a): Alger; Tala-Guizan. Corbière (1889), Orgaz et al.
Records. — Boukhatem et al. (2017): El-Kala National Park, (2013): Blida, forêt de Lièges. Cardot (1904): Environs d’Alger,
Tonga lake watershed. Bouzarea. Corbière (1904): Alger, propiété Braham Réis; Bois de
Boulogne; Environs d’Alger, Birmandréis. Corbière (1904), Jelenc
(1955a): Batna, flanc nord du djebel Tougourth. Corbière (1904,
Comment 1905), Jelenc (1955a): Constantine, djebel Ouach. Litschauer
Doubtfully present in Algeria. The presence of this Tropical (1904): Chiffaschlucht, 200 m; Michelet, 1000 m. Litschauer
(1904), Jelenc (1955a): Col de Tirourda, 1700 m; Philippeville,
montane species (Dierßen 2001) in the Mediterranean Region 50 m; Qued-Kebir bei Blidah, 200 m. Corbière (1905): Envi-
is unlikely and requires verification. rons de Constantine, Le Hamma; Environs de Constantine, Les
Pins. Corbière (1905), Jelenc (1955a): Environs de Constantine,
Menidj. Lapie (1909): Kabylie du Djurjura. Jelenc (1949, 1967):
Brachymenium commutatum var. attenuatum Monts de Tlemcen, forêt de Zariflet, 1150 m. Jelenc (1953):
Aurès, flanc nord du Chélia, 1800 m; Aurès, flanc nord du dje-
Thér. & Trab. bel Pharaoun, 1800 m. Jelenc (1953, 1967): Bèlezma, cédraies
de l’oued Chaba, 1400 m. Jelenc (1955a): Aizer; Babor; Djebel
Records. — Thériot (1931, 1933), Jelenc (1955a): Atakor-n-Aha- Dira; Kabylie du Djurdjura, Michelet, 1000 m; Médéa; Monts
ggar, Ilaman, 2000 m; Tefedest, Ahetes, 1200 m. Frahm (1978): de Constantine, djebel Chettaba. Bizot (1973): Djebel Belezma
Hoggar, Guelta von Imeghra, 1780 m; Hoggar, Tahabort NE Ta- à 15 km à l’ouest de Batna. Ros et al. (1990): Texenna’s wood,
manrasset, 1700 m. 700 m. Boukhatem et al. (2017): El-Kala National Park, Tonga
lake watershed. Common.
Comment
This taxon was described from Algeria (Thériot 1931) and later
reported from other African countries (Ochi 1972; O’Shea Brachythecium albicans (Hedw.) Schimp.
2006; Ros et al. 2013). According to Ros et al. (2007b, 2013),
the taxonomic value of this name is doubtful. Unfortunately Records. — Jelenc (1955a): Kabylie du Djurdjura, Aizer; La
the type could not be traced by Ros et al. (2007b). Bouzaréa. Rare.

CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5) 67

 Ros R. M. et al.

Comment dah, 300 m. Corbière (1905): Environs de Constantine, Les Pins;

Jelenc (1955a) informed that according to Cardot, the iden- Environs de Constantine, route de la Corniche. Corbière (1905),
Jelenc (1955a): Environs de Constantine, djebel Ouach. Camus
tification of the specimen of “La Bouzaréa” is doubtful. (1906), Coppey (1911), Jelenc (1955a): Beni-Ounif, Sahara algé-
rien. Braun-Blanquet & Maire (1924), Jelenc (1955a): Tlemcen.
Jelenc (1955a): Arba; Col des Aït Aïcha; Perrégaux; Sud algérois,
Brachythecium campestre (Müll. Hal.) Schimp. Bou Saâda; Sud algérois, Laghouat. Jelenc (1956, 1967): Sahel de
Koléa. Boukhatem et al. (2017): El-Kala National Park, Tonga lake
Records. — Bescherelle (1882), Jelenc (1955a): Alger. Very rare. watershed. Mazari & Boulaacheb (2023): Setif, Megriss Mountain.
Common.

Comment
The only report of this Austral-montane-arctic species (Dierßen Bryum canariense Brid.
2001) in Algeria is based on a single specimen collected by
Durieu de Maisonneuve. Records. — Bescherelle (1882), Jelenc (1955a): Constantine. Dixon
(1931), Ochi (1972): Taharauet, southern part of Hoggar Mts.,
Central Sahara, 5900 ft. Jelenc (1955a): Hoggar, Fort Motylinsky,
1900 m; Sud oranais; Tiaret. Rare.
Brachythecium rivulare Schimp.

Records. — Feldmann (1947), Jelenc (1955a), Orgaz et al. (2012): Bryum dichotomum Hedw.
Kabylie de Collo, oued Elli Ankou, 500-600 m. Boukhatem et al.
(2017): El-Kala National Park, Tonga lake watershed. Rare.
Bryum bicolor var. denticulatum (Besch.) Jelenc
Bryum bicolor var. macrothecium (Besch.) Jelenc
Brachythecium rutabulum (Hedw.) Schimp.
Bryum bicolor var. tenellum (Besch.) Jelenc
Records. — Bescherelle (1882): Atlas; Kabylie. Bescherelle (1882),
Jelenc (1955a): Bône; Édough; La Calle. Gandoger (1883), Trabut Nomenclature follows Holyoak (2021).
(1888): sine loco. Litschauer (1904), Jelenc (1955a): Fort National,
1000 m; Im Tal des Qued-Kebir bei Blidah, 200 m. Corbière (1905): Records. — Bescherelle (1882): Alger, Fort l’Empereur; remparts,
Environs de Constantine, djebel Ouach; Environs de Constantine, près de la porte Randon. Bescherelle (1882), Camus (1906), Jelenc
Les Pins; Environs de Constantine, Menidj; Environs de Con- (1955a): Oran. Bescherelle (1882), Jelenc (1955a): Alger; Blidah;
stantine, route du Séminarie. Jelenc (1955a): Alger, La Bouzaréa; Bône; Bougie; Gorges de la Chiffa; Philippeville. Corbière (1889):
Constantine; Kabylie du Djurdjura, Akfadou; Kabylie du Djurd- Blida, grèves l’oued Beni-Aza, sommet des Beni-Salah. Julien (1894):
jura, Tizi Ouzou. Boukhatem et al. (2017): El-Kala National Park, Région de Constantine. Corbière (1904): Médéa. Corbière (1905):
Tonga lake watershed. Mazari & Boulaacheb (2023): Setif, Megriss Environs de Constantine, Chettabah; Environs de Constantine,
Mountain. Frequent. djebel Ouach; Environs de Constantine, Moulin Laire; Environs de
Constantine, oued Zied; Environs de Constantine, route de Sétif;
Environs de Constantine, Sidi M’Cid. Braun-Blanquet & Maire
(1924): Tlemcen. Jelenc (1951): Sahel d’Oran, djebel Murdjadjo, forêt
Brachythecium salebrosum de M’sila à Msabia; Sahel d’Arzeu, djebel Borouse. Jelenc (1955a):
(Hoffm. ex F.Weber & D.Mohr) Schimp. Alger; Atlas de Blida, oued el Kebir, 300 m; Aumale; Bône; Djebel
Chenoua; Djebel Ouach; Frais Vallon, 300 m; Kabylie des Babor,
Records. — Bescherelle (1882): Kabylie. Bescherelle (1882), Jelenc cap Aokas; Kabylie du Djurdjura, Bouzgeur; Kabylie du Djurdjura,
(1955a): Constantine, rochers; Tala-Guizan. Jelenc (1955a): Kabylie Fort National; La Calle; La Réghaia; Michelet, 1000 m; Monts de
du Djurdjura, Azerou n‘Tiddjer. Orgaz et al. (2012): Oyan. Boukha- Constantine, djebel Chettaba; Ouarsenis, djebel Doui; Taourirt; Ti-
tem et al. (2017): El-Kala National Park, Tonga lake watershed. Rare. rourda. Jelenc (1955a, 1955b, 1967): Monts de Constantine. Jelenc
(1955a), Ahayoun et al. (2007): Dellys. Jelenc (1956): Golfe de Mo-
staganem, junipéraie de la Macta; Haute Plaine de Sidi-bel-Abbès.
Jelenc (1956, 1967): Massif de la Bouzaréa, forêt de Baïnem; Massif
Bryoerythrophyllum recurvirostrum (Hedw.) P.C.Chen du Dahra, junipéraie du cap ivi. Jelenc (1967): Haute-Plaine de Sidi
Bel Abés, Baudens; Massif du Tessala, pic de Tafaraoui, 650 m. Bizot
Records. — Bescherelle (1882), Jelenc (1955a): Tiaret. Bizot (1973): Ruines de Timgad. Ochi (1973): Algier, Bona, somaische
(1973): Djebel Belezma à 15 km à l’ouest de Batna. Boukhatem Ruine; Mustapha de Alger. Frahm (1978): Béchar. Ahayoun et al.
et al. (2017): El-Kala National Park, Tonga lake watershed. Rare. (2007): sine loco. Ros et al. (1990): Between Mostaganem and Cap Ivi,
Abdelmaimalek-Ramdan beach, 0 m; Cape Falcon, les Andalouses,
30-50 m; Ouled Mimoum; Skikda beach, 0 m. Boukhatem et al.
Bryum argenteum Hedw. (2017): El-Kala National Park, Tonga lake watershed. Very common.

Records. — Bescherelle (1882): Aïn El-Hammam; Hôpital du

Dey; Kabylie; Kasbah; Mustapha. Bescherelle (1882), Jelenc (1955a):
Alger; Blidah; Bou-Khalfa; Constantine, rochers du ravin du Rum- Bryum erythrocarpum complex
mel; La Calle, rochers de grès; Rovigo. Gandoger (1883): sine loco.
Corbière (1889): Blida, Nouvelle-Blida. Julien (1894): Région de Records. — Montagne (1838): Alger. Jelenc (1949, 1967): Massif
Constantine. Corbière (1904): Alger, talus de Telemly. Corbière du Murdjadjo (Oran), ravin de Noiseux. Jelenc (1955a): Afensou;
(1904), Jelenc (1955a): Médéa. Litschauer (1904), Jelenc (1955a): Kabylie du Djurdjura, Fort National; Monts de Tlemcen, forêt
Michelet und Fort National, 1000-1200 m; Qued-Kebir bei Bli- d’Hafir; Tirourda. Jelenc (1967): Djebel Borousse.

68 CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5)

 Revisiting the Bryophyte Checklist of Algeria

Comment Calcidicranella howei (Renauld & Cardot)

The concept of Crundwell & Nyholm (1964) is followed for Bonfim Santos, Fedosov & Jan Kučera
the definition of this complex.
The names reported in Algeria are Bryum erythrocarpum Dicranella howei Renauld & Cardot
Schwägr. and Bryum sanguineum Brid. nom. illeg. incl. spec. prior. Nomenclature follows Fedosov et al. (2023).
Records. — Bescherelle (1882): Alger; Birmandreïs; Coteaux de
Bab-Azoun; Pointe-Pescade; Près Blidah et Boufarik. Bescherelle
Bryum gemmilucens R.Wilczek & Demaret (1882), Jelenc (1955a): Bouzaréah; Maison-Carrée. Corbière (1889):
Blida. Corbière (1904): Alger, talus de la route de Telemly, 80 m.
Records. — Ros et al. (1990): Dellys, 0-40 m. Very rare. Corbière (1905), Jelenc (1955a): Environs de Constantine, bords
du Rummelet de l’oued Bou Merzoug; Environs de Constantine,
Le Hamma. Jelenc (1955a): Atlas de Blida, oued el Kebir, 200 m;
Boufarik; Kabylie du Djurdjura, Tifrit n’Aït el Hadj. Ros et al. (1990):
Bryum gemmiparum De Not. Between Mostaganem and Cap Ivi, Abdelmaimalek-Ramdan beach,
0 m; Texenna’s wood, 700 m. Frequent.
Bryum gemmiparum var. ahhagarense Thér. & Trab.
Nomenclature follows Holyoak (2021).
Calcidicranella varia
Records. — Montagne (1849, 1856), Bescherelle (1882), Tra- (Hedw.) Bonfim Santos, Fedosov & Jan Kučera
but (1913), Jelenc (1955a), Ahayoun et al. (2007): Prope Tiaret.
Bescherelle (1882): Bouïram; Constantine, aux moulins; Kaby- Dicranella varia (Hedw.) Schimp.
lie. Bescherelle (1882), Jelenc (1955a): Alger; Blidah; Col d’Aïn
Kebira; La Calle, pâturages humides; Hoggar, mont Asekrem, Nomenclature follows Fedosov et al. (2023).
2400 m; Milah; Tin Seghin. Bescherelle (1882), Trabut (1914):
Djurdjura. Gandoger (1883): sine loco. Corbière (1904): Alger, Records. — Montagne (1838), Jelenc (1955a): Alger. Gandoger
rochers humides, route de Telemly, 80 m. Corbière (1905): En- (1883), Trabut (1888): sine loco. Jelenc (1951): Haute plaine de
virons de Constantine, route de la Corniche. Corbière (1905), Tlemcen, berges du Chabet bel Abbès, 750 m; Monts de Tlemcen,
Jelenc (1955a): Environs de Constantine, Sidi M’Cid. Thériot forêt de Zariffet, l’oued Zarifète, 1150 m. Jelenc (1951), Jelenc
(1931, 1933): Atakor-n-Ahaggar ad meridiem montis Asekrem, (1967): Sahel d’Oran, djebel Murdjadjo au ravin de Noiseux; Sahel
2400 m. Jelenc (1951, 1967): Monts de Tlemcen, oued en Nechef, d’Arzeu, djebel Borouse. Jelenc (1953, 1967): Babor, flanc nord du
1100 m; Forêt de Zariffet, 1100-1200 m. Jelenc (1955a): Djebel Babor, ravin des Beni Bezez, 1300 m. Jelenc (1954b): Sahel d‘Alger,
Smir; Kabylie du Djurdjura, Bouira; Tiaret, cascade de la Mina. colline du Hamma. Jelenc (1955a): Ben Chicao; Gorges de la Chiffa,
Jelenc (1956, 1967): Massif de la Bouzaréa. Ochi (1972): Cisret. 300-400 m. Jelenc (1956, 1967): Massif de la Bouzaréa, ravin de
Boukhatem et al. (2017): El-Kala National Park, Tonga lake wa- la prise d’eau près Bains-Romanis; Plaine de la Mitidja, à Baba-Ali,
tershed. Common. sur les bords de l’oued Kerma; Sahel d’Alger, ravin du Frais-Vallon.
Jelenc (1966, 1967): Sahel de Zéralda. Jelenc (1967): Boufarik;
Maison-Carrée; Monts et Haute-Plaine de Tlemcen. Boukhatem
et al. (2017): El-Kala National Park, Tonga lake watershed. Frequent.
Bryum kabylicum Trab. in Jelenc nom. inval.

Records. — Jelenc (1955a): Djurdjura. Callicladium imponens

(Hedw.) Hedenäs, Schlesak & D.Quandt
Comment Hypnum imponens Hedw.
Although included in the checklists of Ros et al. (1999, 2013),
the identity of this name seems not to have been revised Nomenclature follows Schlesak et al. (2018).
after having been proposed. Unfortunately, its identity could Records. — Jelenc (1949, 1967): Monts de Tlemcen, djebel el
not be assessed because the type could not be traced (Ros, Béniane, 1050 m. Boukhatem et al. (2017): El-Kala National Park,
unpublished). Tonga lake watershed. Rare.

Bryum radiculosum Brid. Calliergonella cuspidata (Hedw.) Loeske

Records. — Bescherelle (1882): Kabylie. Jelenc (1955a): Kabylie

Records. — Bescherelle (1882): Bab-Azoun; Bouzaréah; Cons- du Djurdjura, Tizi n’Djemaâ; Monts de Constantine. Boukhatem
tantine; Djebel Mansourah; La Chiffa. Bescherelle (1882), Jelenc et al. (2017): El-Kala National Park, Tonga lake watershed. Mazari &
(1955a): La Calle. Bescherelle (1882), Litschauer (1904), Jelenc Boulaacheb (2023): Setif, Megriss Mountain. Rare.
(1955a), Ochi (1972): Alger. Corbière (1904): Maison Carrée près
Alger. Litschauer (1904): Chiffaschlucht und im Tal des Qued-Ke-
bir bei Blidah, 300 m. Jelenc (1955a): La Bouzaréa. Jelenc (1956):
Maison-Carrée, Ecole National d’Agriculture. Bizot (1973): Djebel Campylopus brevipilus Bruch & Schimp.
Belezma à 15 km à l’ouest de Batna; El Golea; Fontaine Chaude à
30 km NE de Batna. Boukhatem et al. (2017): El-Kala National Records. — Bescherelle (1882): sine loco. Jelenc (1955a): Kabylie
Park, Tonga lake watershed. Mazari & Boulaacheb (2023): Setif, du Djurdjura, Aït Idjer. Frahm (1985): Kabylie. Boukhatem et al.
Megriss Mountain. Frequent. (2017): El-Kala National Park, Tonga lake watershed. Rare.

CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5) 69

 Ros R. M. et al.

Campylopus flexuosus (Hedw.) Brid. D’Oran à Mécheria. Corbière (1904), Jelenc (1955a): Orléansville,
berges du Chélif, 130 m. Jelenc (1955a): Atlas de Blida; Oran. Ma-
Records. — Bescherelle (1882), Jelenc (1955a): Djebel Aurès. zari & Boulaacheb (2023): Setif, Megriss Mountain. Rare.
Boukhatem et al. (2017): El-Kala National Park, Tonga lake wa-
tershed. Rare.

Cinclidotus aquaticus (Hedw.) Bruch & Schimp.

Campylopus introflexus (Hedw.) Brid.
Records. — Bescherelle (1882), Jelenc (1955a): Oued En-Nekob,
Records. — Boukhatem et al. (2017): El-Kala National Park, au pied du djebel Tababor. Jelenc (1955a): Kabylie des Babor; Kabylie
Tonga lake watershed. Very rare. du Djurdjura, Ansor Aberkane. Jelenc (1966, 1967): Haute-Plaine
de Tlemcen, ruisseau près de l’Ain Defla, 650 m. Ahayoun et al.
(2007): Djurdjura. Boukhatem et al. (2017): El-Kala National Park,
Comment Tonga lake watershed. Rare.
Jelenc (1955a) reported this species based on the previous one
of Bescherelle (1882) from La Calle as Campylopus polytrichoides
De Not. considering both names as synonyms. Nevertheless, Cinclidotus fontinaloides (Hedw.) P.Beauv.
posterior taxonomical studies by Gradstein & Sipman (1978)
differentiate clearly both species and confirmed that at that Records. — Bescherelle (1882), Jelenc (1955a): Djurdjura. Monts
time the reports in northern Africa correspond to C. pilifer. de Tlemcen, sources de la Tafna. Jelenc (1951, 1967): Monts de
Tlemcen, oued Mefrouche. Boukhatem et al. (2017): El-Kala Na-
tional Park, Tonga lake watershed. Rare.

Campylopus pilifer Brid.

Cirriphyllum crassinervium (Taylor) Loeske & M.Fleisch.
Records. — Bescherelle (1882): La Calle. Very rare.
Records. — Bescherelle (1882): Bône. Jelenc (1949, 1967): Monts
Comment de Tlemcen, forêt de Zariflet, 1100 m. Boukhatem et al. (2017):
El-Kala National Park, Tonga lake watershed. Rare.
It was not listed for Algeria by Ros et al. (1999, 2013) based
on the criterium of Jelenc (1955a) (see comment in the for-
mer species). Cirriphyllum piliferum (Hedw.) Grout

Records. — Lapie (1909), Jelenc (1955a): Kabylie du Djurdjura,

Campylostelium strictum Solms Aizer. Very rare.

Records. — Feldmann (1947), Jelenc (1955a): Kabylie de Collo,

Taoulel. Boukhatem et al. (2017): El-Kala National Park, Tonga Codonoblepharon forsteri (Dicks.) Goffinet
lake watershed. Rare.
Zygodon forsteri (Dicks.) Mitt.
Nomenclature follows Goffinet et al. (2004) and Hodgetts et al.
Ceratodon purpureus (Hedw.) Brid. (2020).
Records. — Bescherelle (1882), Jelenc (1955a): Kabylie du Djurd- Records. — Jones (1956), Jelenc (1967): Yakouren (nr. Azazga),
jura, Tizi Ouzou. Lapie (1909): Kabylie du Djurjura. Feldmann Kabylie Mountains. Very rare.
(1947): Kabylie de Collo, vieilles souches de Chêne-Liège, 800 m,
aux environs de Bessombourg. Jelenc (1955a): Kabylie du Djurd-
jura, Aït Ouaban; Kabylie du Djurdjura, Dra Inguel. Jelenc (1956, Coscinodon cribosus (Hedw.) Spruce
1967): Sahel d’Alger, près de Zèralda. Boukhatem et al. (2017):
El-Kala National Park, Tonga lake watershed. Rare. Records. — Jelenc (1955a): Gorges de la Chiffa, 300-400 m.
Very rare.

Ceratodon purpureus subsp. stenocarpus Cratoneuron filicinum (Hedw.) Spruce

(Bruch & Schimp. ex Müll. Hal.) Dixon
Records. — Bescherelle (1882): Ehna-Bord; Ravins de l‘oued
Records. — Jelenc (1955a): Kabylie de Collo, Bessombourg; Ou- Sidi-el-Kebir. Bescherelle (1882), Jelenc (1955a): Atlas de Blida,
arsenis, Téniet el Had. Mazari & Boulaacheb (2023): Setif, Megriss Aïn Talazid; Djurjura; Gorges de la Chiffa; Massif de Tablat,
Mountain. Rare. Sakamody; Monts de Constantine. Corbière (1904): Tlemcen,
parois de la dernière vasque de la cascade. Corbière (1904), Jelenc
(1955a): Batna, parois d’une source calcaire, flanc nord du djebel
Cheilothela chloropus (Brid.) Broth. Tougourth, 1300 m; Batna, rochers sur le flanc sud du djebel
Tougourth, 1300 m. Litschauer (1904): Qued-Kebir bei Blidah,
Records. — Bescherelle (1882): La Galite; Oran, djebel Santo. 300 m. Jelenc (1949, 1967): Monts de Tlemcen, cirque d’El
Bescherelle (1882), Jelenc (1955a): Philippeville. Trabut (1887a): Ourit, 1100 m; Monts de Tlemcen, Plateau travertineux de Lalla

70 CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5)

 Revisiting the Bryophyte Checklist of Algeria

Setti. Jelenc (1953): Bèlezma, pentes sud-est du djebel Touggour, Comment

1700 m. Jelenc (1953, 1967): Massif de Aurès, flanc nord du Crossidium squamiferum var. brevisetum (Besch.) Paris was
Chélia, 1900 m; Massif de Aurès, ravin de la maison forestière
du Chélia, 1500 m; Massif de l’Aurès, forêt de Sgag, 1650 m;
described by Bescherelle (1882) as Barbula membranifolia f.
Monts du Hodna, Maadid, maison forestière de Mahali, 1300 m. breviseta Besch. from three localities (Constantine, Ravin du
Jelenc (1955a): Atlas de Blida, oued el Kebir, 200 m; Djurdjura, Rummel and Laghouat). According to Ros et al. (2013) its
Lalla Khadidja; Monts de Tlemcen, cascade de Mazer. Jelenc identity has not been revised since. Unfortunately the type
(1955b, 1967): Monts de Hodna au massif du Bou Thaleb à At- could not be traced (Ros, unpublished). Probably it is within
tafi, 1400 m. Jelenc (1967): Assif du Bélézma, djebel Touggour,
1700 m. Boukhatem et al. (2017): El-Kala National Park, Tonga the morphological variability of the species, as it was stated
lake watershed. Common. for var. pottiodeum by Cano et al. (1993) and Hodgetts et al.
(2020).

Crossidium aberrans Holz. & E.B.Bartram

Crossidium squamiferum var. pottioideum
Records. — Delgadillo (1975): Oran. Cano (1992): El Kantara. (De Not.) Mönk.
Boukhatem et al. (2017): El-Kala National Park, Tonga lake wa-
tershed. Rare. Records. — Corbière (1904), Jelenc (1955a): Tiaret, Hauts-Plateaux,
Douar Torrich, 1000 m. Corbière (1905): Environs de Constantine,
sur les travertins du Mansourah. Jelenc (1955a): Constantine; Oran;
Crossidium crassinervium (De Not.) Jur. Sud algérois, Bou Saâda; Sud algérois, Laghouat. Jelenc (1967):
Monts de Tlemcen, djebel el Béniane, 950 m. Delgadillo (1975):
Records. — Bescherelle (1882), Jelenc (1955a): Birmandreïs, bord El Guerah. Rare.
des chemins; Oued Sidi-Tifour, près de Macta, sud de la province
d’Oran. Bescherelle (1882), Jelenc (1955a), Delgadillo (1975):
Alger. Trabut (1887a): D’Oran à Mécheria. Corbière (1905): En-
virons de Constantine, Les Pins. Camus (1906), Coppey (1911), Cryphaea heteromalla (Hedw.) D.Mohr
Jelenc (1955a): Beni-Ounif, Sahara algérien; Ravin de Ben Zireg,
dans l’extrème-Sud Oranais. Camus (1906), Jelenc (1955a): envi- Records. — Bescherelle (1882): Philippeville. Bescherelle (1882),
rons de Tlemcen. Jelenc (1955a): Chebka, oued Mredjar au Nord Jelenc (1955a): Alger, sur les lentisques, les oliviers; Djebel Thaya;
de Ghardaïa; Constantine; Hauts-plateaux oranais; Orléansville; Kabylie du Djurdjura, Akfadou; La Calle. Trabut (1888): sine
Saoula; Sud constantinois, Biskra. Jelenc (1966, 1967): Sahel d’Arzeu loco. Corbière (1904): La Calle, fôret de l’oued Mellah. Corbière
à Renan. Boukhatem et al. (2017): El-Kala National Park, Tonga (1905), Jelenc (1955a, 1966, 1967): Monts de Constantine, dje-
lake watershed. Frequent. bel Ouach, 1000 m. Jelenc (1955a): Castiglione; Gorges de la
Chiffa; Kabylie du Djurdjura, Tifrit; Khodjaberry; Koléa. Jelenc
(1956, 1967): Atlas Mitidjien, col des Deux-Bassins, 850 m.
Boukhatem et al. (2017): El-Kala National Park, Tonga lake
Crossidium laevipilum Thér. & Trab. watershed. Frequent.

Records. — Thériot (1931, 1933), Jelenc (1955a): Ghardaïa, Mzab.

Jelenc (1955a): Sud algérois, Laghouat. Boukhatem et al. (2017): Ctenidium molluscum (Hedw.) Mitt.
El-Kala National Park, Tonga lake watershed. Rare.
Records. — Bescherelle (1882): Oued Djer; Sud d‘Oran. Besche-
relle (1882), Jelenc (1955a): Édough; Gorges de la Chiffa; Oued
Crossidium squamiferum (Viv.) Jur. var. squamiferum Merdja. Trabut (1887a): D‘Oran à Mécheria. Jelenc (1953, 1967):
Kabylie des Babor, crête du Tababor, 1950 m. Jelenc (1955a): Atlas
Records. — Bescherelle (1882): Constantine; Couteaux de Bab- de Blida; Crêtes du Djurdjura, Aït Ouaban; Hauts-plateaux ora-
Azoun. Bescherelle (1882), Jelenc (1955a): Alger; Bougie; Bou-Sâada; nais. Boukhatem et al. (2017): El-Kala National Park, Tonga lake
Saoula. Corbière (1889), Jelenc (1955a): Blida, oued Sidi-el-Kebir. watershed. Frequent.
Cardot (1904), Maire (1916): province d’Oran, Djebel Morghad,
1950 m, 2105 m. Corbière (1904): Alger, talus à Telemly. Cor-
bière (1904), Jelenc (1955a): Nador de Médéa. Corbière (1905):
Environs de Constantine, Chettabah. Corbière (1905), Jelenc Cynodontium bruntonii (Sm.) Bruch & Schimp.
(1955a): Environs de Constantine, Sidi M’Cid. Camus (1906):
Oran; environs de Tlemcen. Camus (1906), Coppey (1911), Jelenc Records. — Feldmann (1947), Jelenc (1955a): Kabylie de Collo,
(1955a): Beni-Ounif, Sahara algérien. Jelenc (1949): Haute plainte sommet du djebel Goufi, versant Nord, 1000 m. Boukhatem et al.
de Tlemcen, berges d l’oued Sennoun, 350 m, Monts de Tlemcen, (2017): El-Kala National Park, Tonga lake watershed. Rare.
pentes du djebel el Béniane, 950 m. Jelenc (1954b, 1966, 1967):
Atlas Mitidjien, gorges de l’oued Harrach à Hammam Mélouane,
150 m. Jelenc (1954b, 1967): Massif du Bou Zegza, gorges de Ked- Dialytrichia mucronata (Brid.) Broth.
dara, 400 m. Jelenc (1955a): Aumale; El Kantara, 1500 m; Gorges
de la Chiffa; La Bouzaréa; La Réghaia; Monts de Constantine, Records. — Bescherelle (1882): Petit Atlas; sine loco. Corbière
djebel Ouach; Monts de Tlemcen, Tlemcen, forêt d’Hafir; Sud (1905): Environs de Constantine, ravin du Rummel; Environs de
constantinois, Biskra; Téniet Zebboudj. Jelenc (1967): Haute-Plaine Constantine, Sidi M’Cid. Jelenc (1954b): Massif du Bou Zegza,
de Tlemcen. Delgadillo (1975): Atlab, Ben Zireg. Frahm (1978): gorges de Keddara, 400 m. Jelenc (1955a): Alger; Atlas de Blida;
Laghouat. Boukhatem et al. (2017): El-Kala National Park, Tonga Constantine; Djurdjura. Boukhatem et al. (2017): El-Kala National
lake watershed. Common. Park, Tonga lake watershed. Rare.

CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5) 71

 Ros R. M. et al.

Dicranella heteromalla (Hedw.) Schimp. Maire (1924): Tlemcen. Jelenc (1951, 1955a, 1967): Haute plaine
de Tlemcen, vallée de l’oued el Horra. Jelenc (1955a): Atlas de
Records. — Jelenc (1966): Alluvions d’un affluent de l’oued Blida, oued el Kebir, 300 m; Col de Tirourda, 1700 m; Kabylie des
Ouadallah, 1100 m. Jelenc (1966, 1967): Monts de Tlemcen, dans Babor, cap Aokas; Laghouat; Maison-Carrée. Bizot (1973): Djebel
la forêt de Zariffet. Boukhatem et al. (2017): El-Kala National Park, Belezma à 15 km à l’ouest de Batna. Ros et al. (1990): Cape Fal-
Tonga lake watershed. Rare. con, les Andalouses, 30-50 m; Hammam-Riga, 500 m. Ros et al.
(1990), Jiménez et al. (2005b): Between Mostaganem and Cap
Ivi, Abdelmaimalek-Ramdan beach, 0 m. Jiménez et al. (2005b):
Algiers, route de la Mouzaia; Bosque Texenna; Oued Rhiou; St.
Dicranoweisia cirrata (Hedw.) Lindb. Eugène. Jiménez (2006): Algiers, Sidi Madani. Boukhatem et al.
(2017): El-Kala National Park, Tonga lake watershed. Common.
Records. — Bescherelle (1882), Jelenc (1955a): Djebel Édough,
forêt de Buïn-Si; La Calle, vallée d’Ain-Crier. Jelenc (1953, 1967):
Aurès, flanc nord du djebel Pharaoun, 1800 m. Jelenc (1955a):
Camp de la Santé; Kabylie du Djurdjura, Fort National; Ouarsenis, *Didymodon giganteus (Funck) Jur.
Téniet el Had; Taourirt. Bizot (1973): Djebel Belezma à 15 km à
l’ouest de Batna. Boukhatem et al. (2017): El-Kala National Park, Barbula nivalis (Spruce) A.Jaeger
Tonga lake watershed. Rare.
Records. — Julien (1894): Région de Constantine.

Comment
Dicranum scoparium Hedw. Doubtfully present in Algeria. T h is report was published as
Records. — Bescherelle (1882): Djurjura. Bescherelle (1882), Jelenc
Barbula nivalis (without the author’s name) but indicating
(1955a): Tala-Guizan. Jelenc (1955a): Forêt d’Akfadou; Kabylie du that it was one of the most common mosses in the region. As
Djurdjura, Tiruourda. Boukhatem et al. (2017): El-Kala National the identity of the type of the basionym of this name (Tor-
Park, Tonga lake watershed. Mazari & Boulaacheb (2023): Setif, tula vinealis var. nivalis Spruce) is Didymodon gianteus and it
Megriss Mountain. Rare. is a rare species in the Mediterranean Region, it is doubtful
to which species Julien referred. It was not included in the
previous checklists of Ros et al. (1999, 2013).
Dicranum tauricum Sapjegin

Records. — Jelenc (1955a, 1955b, 1967): Massif de l’Aurès au

djebel Chélia. Very rare. Didymodon insulanus (De Not.) M.O.Hill

Records. — Montagne (1849, 1856): Tournit. Bescherelle (1882):

Didymodon acutus (Brid.) K.Saito Toumiet, sur les bords du ruiseau, au bas de la colline. Bescherelle
(1882), Jelenc (1955a): Alger, près Médéah; Bône; Col des Toumi-
Records. — Bescherelle (1882): Birmandreïs. Bescherelle (1882), ettes; Constantine; Djurdjura. Corbière (1889), Jelenc (1955a):
Jelenc (1955a): Alger; Blidah; Bouzaréah. Corbière (1904), Jelenc Blida. Corbière (1904), Jelenc (1955a): Médéa. Corbière (1904,
(1955a): Sétif, 1100 m. Corbière (1905): Environs de Constan- 1905), Jelenc (1955a): Constantine, bois du djebel Ouasch, talus
tine, Les Pins. Jelenc (1955a): Arba; Atlas de Blida, oued el Kebir, d’un fossé, 1000 m. Jelenc (1955a): Aumale; Kabylie des Babor,
300 m; Constantine; Massif du Chénoua; Oran; Pointe-Pescade. cap Aokas; Kabylie du Djurdjura, Fort National; Tell oranais. Jelenc
Jelenc (1955a), Ros et al. (1990): Gorges de la Chiffa. Ros et al. (1956): Kabylie du Djurdjura, massif de l’Arkafadou à la Fontaine
(1990): Oued Rhiou; Ouled Mimoum. Jiménez (2006): Tizi-Ou- des Ifs. Jelenc (1967): Atlas Mitidjien, pic Mouzaïa; Haute-Plaine
zou. Boukhatem et al. (2017): El-Kala National Park, Tonga lake de Tlemcen, Chabet-bel-Abbès, 750 m et oued Sennoun; Kabylie
watershed. Frequent. du Djurdjura, Akfadou, 1200 m; Monts de Tlemcen, forêt de Tes-
sera Mramet, 1400 m. Bizot (1973): Ruines de Timgad. Ros et al.
(1990): Texenna’s wood, 700 m. Ros et al. (1990), Jiménez (2006):
Didymodon australasiae (Hook. & Grev.) R.H.Zander Chiffa’s Gorge. Cano et al. (2000): sine loco. Boukhatem et al. (2017):
El-Kala National Park, Tonga lake watershed. Mazari & Boulaacheb
(2023): Setif, Megriss Mountain. Common.
Records. — Ros & Guerra (1987a), Jiménez et al. (2005a), Jiménez
(2006): Constantine, El Kantara. Boukhatem et al. (2017): El-Kala
National Park, Tonga lake watershed. Rare.
Didymodon luridus Hornsch.
Didymodon fallax (Hedw.) R.H.Zander Records. — Bescherelle (1882), Jelenc (1955a): Alger. Corbière
(1889), Jiménez (2006): Blidah. Corbière (1904), Jelenc (1955a):
Records. — Bescherelle (1882), Jelenc (1955a): Alger; Bône; Kab- Médéa; Sétif, 1085 m. Corbière (1905): Environs de Constantine,
ylie du Djurdjura, Tizi Ouzou, 1500 m; Tiaret. Bescherelle (1882), bords de l’oued Bou Merzoug; Environs de Constantine, Le Hamma;
Corbière (1889), Jiménez et al. (2005b): Blidah. Gandoger (1883): Environs de Constantine, Menidj; Environs de Constantine, Moulin
sine loco. Corbière (1904), Jelenc (1955a): Médéa; Sétif, 1100 m. Laire; Environs de Constantine, route de la Corniche; Environs de
Corbière (1905): Environs de Constantine, Bellevue; Environs de Constantine, Sidi M’Cid. Braun-Blanquet & Maire (1924), Jelenc
Constantine, Les Pins; Environs de Constantine, Moulin Laire; (1955a): Tlemcen. Jelenc (1955a): Atlas de Blida, oued el Kebir,
Environs de Constantine, route de Sétif; Environs de Constan- 200 m; Bône; Constantine; Djebel Chenoua; Gorges de la Chiffa,
tine, Sidi M’Cid; Environs de Constantine, tuileries du Rummel. 300-400 m; Orléansville; Philippeville. Ros et al. (1990): Oued
Corbière (1905), Jelenc (1955a): Constantine. Braun-Blanquet & Rhiou. Frequent.

72 CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5)

 Revisiting the Bryophyte Checklist of Algeria

Didymodon nicholsonii Culm. Didymodon tophaceus subsp. sicculus

(M.J.Cano, Ros, García-Zamora & J.Guerra) Jan Kučera
Records. — Jiménez et al. (2003), Jiménez (2006): sine loco. Very rare.
Didymodon sicculus M.J.Cano, Ros, García-Zamora & J.Guerra
Nomenclature follows Kučera et al. (2018).
Didymodon rigidulus Hedw.
Records. — Jiménez et al. (2004), Jiménez (2006): Constantine,
Records. — Montagne (1838), Bescherelle (1882), Jelenc (1955a), El Kantara. Very rare.
Ros & Cano (2008): Alger. Jelenc (1955a): Atlas de Blida, oued el
Kebir; Kabylie du Djurdjura, Fort National, 1000 m. Jelenc (1955a),
Jiménez (2006), Ros & Cano (2008): Gorges de la Chiffa. Bizot Didymodon vinealis (Brid.) R.H.Zander
(1973): Fontaine Chaude à 30 km NE de Batna; Ruines de Tim-
gad. Ros & Guerra (1987a): El Kantara. Boukhatem et al. (2017): Records. — Bescherelle (1882): Birmandreïs; Bouzaréah; Douéra;
El-Kala National Park, Tonga lake watershed. Rare. Kabylie. Bescherelle (1882), Corbière (1904), Jelenc (1955a): Médéah.
Bescherelle (1882), Corbière (1905), Jelenc (1955a, 1967): Constan-
tine. Bescherelle (1882), Jelenc (1955a): Alger; Mascara; Philippeville.
Bescherelle (1882), Jelenc (1955a), Jiménez (2006): Batna, Minas
Didymodon tophaceus (Brid.) Lisa subsp. tophaceus de Tringad. Trabut (1887a): D’Oran à Mécheria, Tell. Julien (1894):
Région de Constantine. Corbière (1904), Jelenc (1955a): Gorges
Records. — Bescherelle (1882): Bab-Azoun; Constantine, berges de la Chiffa, ruisseau des Singes; Orléansville, bois de la Pépinière.
du Mansourah; Mustapha. Bescherelle (1882), Corbière (1889), Corbière (1905): Environs de Constantine, Les Pins; Environs de
Jelenc (1955a): Blida, Gorges de la Chiffa. Bescherelle (1882), Jelenc Constantine, Moulin Laire; Environs de Constantine, route de la
(1955a): Alger, bords des ruisseaux; Oran; Tiaret. Trabut (1887a): Corniche; Environs de Constantine, route du Hamma; Environs
D’Oran àa Mécheria. Corbière (1905): Environs de Constantine, de Constantine, tuileries et ravin du Rummel. Jelenc (1949): Haute
Arcades romaines; Environs de Constantine, Le Hamma; Envi- plaine de Tlemcen, Berges de l’oued Sennoun, 350 m; Haute plaine
rons de Constantine, Les Pins; Environs de Constantine, Moulin de Tlemcen, Berges du chabet bel Abbès, 750 m; Monts de Tlemcen,
Laire; Environs de Constantine, route de la Corniche; Environs forêt de Tessera Mramet, 1400 m. Jelenc (1949, 1967): Vallèe de
de Constantine, Salah Bey; Environs de Constantine, Sidi M’Cid. l’Isser, lieu dit “El Kasbat” près de Lamoricière, vers 500 m. Jelenc
Corbière (1905), Jiménez (2006): Environs de Constantine, route (1955a): Azéba; Kabylie du Djurdjura, Azerou ou Gougan, 1700 m;
de Sétif. Thériot (1931, 1933), Jelenc (1955a): Hoggar, Idelès; Kabylie du Djurdjura, Fort National; Pointe-Pescade. Jelenc (1956):
Hoggar, Imarera, 2000 m; Hoggar, Issekkarassen, 2100 m; Hog- Atlas Mitidjien, pic Mouzaïa. Ros & Guerra (1987a): El Kantara.
gar, oued Haman; Hoggar, oued Tarouda; Hoggar, Tassili-n-Ajjer, Ros et al. (1990): Chiffa’s Gorge; Texenna’s wood, 700 m. Mazari &
Amgid. Jelenc (1951, 1967): Monts de Tlemcen, plateau de Lalla Boulaacheb (2023): Setif, Megriss Mountain. Common.
Setti, 1000 m. Jelenc (1953): Babor, flanc nord du Babor, 1300 m;
Bèlezma, flanc sud-est du djebel Touggour, 1700 m. Jelenc (1955a):
Bérard; Bône; Hoggar, Tamanghasset; Kabylie du Djurdjura, col de Distichium capillaceum (Hedw.) Bruch & Schimp.
Tirourda, 1700 m; Kabylie du Djurdjura, Grotte de Tifrit; Monts
de Tlemcen, Mazer; Monts de Tlemcen, Sebdou; Sous, Abouda.
Jelenc (1955a), Jiménez et al. (2005b): Constantine. Collenot et al. Records. — Bescherelle (1882): Djurjura. Bescherelle (1882),
(1960): Tassili N’Aijer, Afaïtalèsse. Jelenc (1967): Aurès, Chélia, Jelenc (1955a): Kabylie du Djurdjura, col de Tirourda; Kabylie des
1800 m; Aurès, forêt de Sgag, 1650 m; Bèlezma, djebel Refaa, Babors, djebel Tababor. Jelenc (1955a): Kabylie du Djurdjura, Aït
2100 m; Djurdjura, Aizer, 1750 m. Frahm (1978): Béchar; Hoggar, Daoud; Kabylie du Djurdjura, forêt d’Akfadou; Ouarsenis. Rare.
Guelta von Imeghra, 1780 m; Hoggar, Gueltas von Issakkarassene,
1800 m. Jiménez et al. (2005b): Hoggar Gebirge, Gueltas von Is-
sakarassene, Tamanghasset; Oasis de Bou Saäda; Philippeville; St. Ditrichum pusillum (Hedw.) Hampe
Eugène; Tizi-Ouzou. Very common.
Records. — Bescherelle (1882), Jelenc (1955a): La Calle. Very rare.

Comment
Three infraspecific taxa were reported from Algeria whose
identity has not been revised after its description (Ros et al. Ditrichum subulatum Hampe
2013): Didymodon tophaceus var. brevicaule (Schimp.) Bou- Records. — Bescherelle (1882): Kabylie. Bescherelle (1882),
lay, reported by Bescherelle (1882) as Trichostomum topha- Jelenc (1955a): La Calle; Philippeville. Corbière (1889), Jelenc
ceum var. brevicaule, and later compiled by Jelenc (1955a) (1955a): Blida, aux Chênes-Lièges, bord d‘un ruisselet, 600 m.
from Alger, le long de la route de Saint-Eugène; Didymodon Jelenc (1954b, 1955a, 1967): Massif de l‘Akfadou, sources sous les
tophaceus fo. acutifolius (Schimp.) Zodda, reported by Jelenc Zéens. Jelenc (1955a): Bône; Édough; Kabylie du Djurdjura, Aït
Daoud; La Réghaia. Rare.
(1955a) from Constantine and Gorges de la Chiffa, ruisseau
des Singes; and Didymodon tophaceus fo. recurvifolius (Bou-
lay) Willd., reported by Jelenc (1951, 1967) from Monts
de Tlemcen, Plateau de Lalla Setti, 1000 m, and by Jelenc Drepanocladus aduncus (Hedw.) Warnst.
(1955a) from Constantine and Monts de Tlemcen, Beni
Records. — Bescherelle (1882), Jelenc (1955a): Kabylie du Djurd-
Snous. Due to the notable morphological variability of the jura, Tala Ghizan; Kabylie du Djurdjura, Igoulmimen. Corbière
species (Jiménez 2006), most probably these names do not (1904): Constantine, bords asséchés de l’étang supérieur du djebel
have any taxonomical value. Ouasch, 1180 m. Jelenc (1955a): Atlas de Blida, oued el Kebir.

CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5) 73

 Ros R. M. et al.

Constantine; Gorges de la Chiffa; Kabylie du Djurdjura, Aït Idjer; oranais, Beni-Ounif, Sahara algérien. Feldmann (1947): Kabylie de
Kabylie du Djurdjura, Akfadou; Kabylie du Djurdjura, Akfadou Collo, Maison forestière de Bou Noghra, 600 m. Jelenc (1955a):
au lac Noir, 1430 m; Kabylie du Djurdjura, Tirourda; La Réghaia; Kabylie du Djurdjura, Fort National; Kabylie du Djurdjura, pic
Maison-Carrée; Monts de Constantine. Boukhatem et al. (2017): Belloua. Frequent.
El-Kala National Park, Tonga lake watershed. Mazari & Boulaacheb
(2023): Setif, Megriss Mountain. Frequent.
Entosthodon commutatus Durieu & Mont.
Comment
Drepanocladus aduncus var. laxus Schimp. ex Jelenc nom. inval. Records. — Durieu de Maisonneuve (1849), Jelenc (1955b), Bru-
gués et al. (2010): sine loco. Montagne (1849, 1856), Bescherelle
was reported from Algeria (Constantine) by Jelenc (1955a ). (1882), Jelenc (1955a): in collibus Babazoum dictis prope Alger.
According to Ros et al. (2013), its identity is difficult to assess Bescherelle (1882): Mustapha. Bescherelle (1882), Jelenc (1955a):
because in his taxonomic study Żarnowiec (2001), was not Édough. Trabut (1922): Beni Oumi de Figuig. Trabut (1927), Jelenc
able to locate the type. (1955a): Sud algérois, entre El Goléa et In Salah; Sud oranais, Beni
Ounif. Thériot (1931, 1933), Jelenc (1955a): Hoggar, oued Haman,
2000 m; Hoggar, oued Tarouda, 2100 m; Hoggar, Tassili-n-Ajjer,
Amgid. Jelenc (1955a): Hoggar, oued Tamanghasset en aval de Fort
Encalypta trachymitria Ripart Laperrine, 1350 m; Mouydir, Tahount Arak. Frequent.

Encalypta rhaptocarpa var. leptodon Lindb.

Entosthodon convexus (Spruce) Brugués
Nomenclature follows Fedosov (2012).
Records. — Jelenc (1955a): Dra el Arba; Kabylie des Babor, cap Records. — Bescherelle (1882), Jelenc (1955a): Alger; Bône; Co-
Aokas. Rare. teau de Caddous. Corbière (1904), Jelenc (1955a): Orléansville,
bois de la Pépinière. Jelenc (1954b, 1967): Massif du Bou Zezga,
gorges de Keddara, 400 m. Jelenc (1955a): Kabylie du Djurdjura,
Encalypta vulgaris Hedw. Fort National; Monts de Tlemcen, forêt d’Hafir. Jelenc (1967):
Boghar, djebel Krellala, 1000 m. Boukhatem et al. (2017): El-Kala
Records. — Bescherelle (1882): Atlas; Batna; Blidah; Constantine, National Park, Tonga lake watershed. Rare.
rochers du Chatabah; Tlemcen. Bescherelle (1882), Jelenc (1955a):
Alger; Massif de la Mahouna, Serdj el Aouda; Monts de Tlemcen,
Mansourah; Sud algérois, Guelt es Stel; Zaccar de Miliana. Corbière
(1889): Blida, bords d’un ruisselet, aux Chênes-Lièges; Blida, grèves Entosthodon duriaei Mont.
de l’oued Beni-Aza. Julien (1894): Région de Constantine. Cardot
(1904): Province d’Oran, Djebel Morghad, 1950 m. Corbière (1904): Records. — Durieu de Maisonneuve (1849): sine loco. Bescherelle
Médéa. Corbière (1904), Jelenc (1955a): Batna, rochers, flanc sud (1882): Alger, pâturages et prairies de l’Agha; Constantine, parois
du djebel Tougourth, 1600 m; Sétif. Corbière (1905): Environs de humides de meules de moulins, rochers calcaires près de la chute
Constantine. Camus (1906), Jelenc (1955a): Oran. Thériot (1931, du Rummel. Bescherelle (1882), Corbière (1904): Alger. Besche-
1933), Jelenc (1955a), Cano et al. (2000): Hoggar, au dessus de relle (1882), Jelenc (1955a): Tlemcen, rochers ombragés, bords
l’oued In-Fergan, 2300 m. Maire (1936): Iles Habibas. Jelenc (1951, des grottes du Mansourah. Battandier & Trabut (1886), Trabut
1967): Massif de Tessala, pic Tafaraoui, 650 m. Jelenc (1954b, 1967): (1887b), Corbière (1904), Brugués (1998): Mustapha. Corbière
Massif du Bou Zegza, gorges de Keddara, 400 m. Jelenc (1955a): (1905), Jelenc (1955a): Environs de Constantine, ravin de Cons-
Aïn Sfa; Aït Khalfoun; Atlas de Blida; Atlas de Blida, oued el Kebir, tantine. Lapie (1909): Kabylie du Djurjura. Jelenc (1955a): Alger,
300 m; Azéba; Bône; Djebel Doui; Djebel Tenouchfi; El Kantara; El Biar; Alger, La Bouzaréa; Djurdjura. Jelenc (1956): Environs d’El
Gorges de la Chiffa; Hauts-plateaux oranais; Kabylie du Djurdjura, Biar; Ravin du Frais-Vallon; Sahel d’Alger, au Bois de Boulogne.
Bouzgeur; Kabylie du Djurdjura, Fort National, 1000 m; Monts Jelenc (1955a), Brugués (1998): Castiglione; Monts de Tlemcen,
de Constantine; Monts des Ksour, djebel Mzi; Nador de Médéa; Mazer. Boukhatem et al. (2017): El-Kala National Park, Tonga lake
Tirourda. Jelenc (1956, 1967): Atlas Miridjien, Meurdja, 1200 m. watershed. Frequent.
Jelenc (1966): Dra Ferchet el Arbi, oued Kamza au djebel Touilila,
1000 m. Jelenc (1966, 1967): Zahrez Chergui, 1000 m. Boukhatem
et al. (2017): El-Kala National Park, Tonga lake watershed. Maza- Entosthodon fascicularis (Hedw.) Müll. Hal.
ri & Boulaacheb (2023): Setif, Megriss Mountain. Very common.
Records. — Bescherelle (1882), Jelenc (1955a): Bône; Philip-
peville. Jelenc (1955a): Arba; Frais Vallon, 300 m; La Bouzaréa;
Comment Maison-Carrée. Rare.
Encalypta vulgaris var. gymnostoma Besch. nom. nud. was
reported from Batna by Jelenc (1955a). According to Ros
et al. (2013) its identity has not been revised since. Entosthodon hungaricus (Boros) Loeske

Records. — Bescherelle (1882), Trabut (1922, 1927), Jelenc

(1955a), Ros & Cano (2008): Const. Sahara Mehounech. Very rare.
Entosthodon attenuatus (Dicks.) Bryhn

Records. — Bescherelle (1882): Kabylie. Bescherelle (1882), Entosthodon muhlenbergii (Turner) Fife
Jelenc (1955a): Alger; Bône; Bouzaréah; La Calle; Philippeville;
Stora. Corbière (1905), Jelenc (1955a): Environs de Constantine, Records. — Montagne (1838), Bescherelle (1882), Corbière
Sidi M’Cid. Camus (1906), Coppey (1911), Jelenc (1955a): Sud (1904), Jelenc (1955a): Alger. Bescherelle (1882), Jelenc (1955a):

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 Revisiting the Bryophyte Checklist of Algeria

Batna; Bône; Kabylie du Djurdjura, Bougie; Mascara; Philippeville; Bescherelle (1882), Jelenc (1955a): Kabylie des Babors, djebel
Tiaret. Trabut (1887a): D’Oran à Mécheria. Corbière (1889): Blida. Tababor; Tiaret. Brongniart (1882): Blidah, cascade. Bescherel-
Julien (1894): Région de Constantine. Corbière (1904): Kabylie, le (1882), Camus (1906), Jelenc (1955a), Cano et al. (2000):
Fort National, 1000 m. Corbière (1904), Jelenc (1955a): Médéa. Tlemcen. Gandoger (1883): sine loco. Trabut (1887a): D‘Oran
Corbière (1905): Environs de Constantine, M’Cid; Environs de à Mécheria. Corbière (1889), Corbière (1904), Jelenc (1955a):
Constantine, route de Sétif. Lapie (1909): Kabylie du Djurjura. Jelenc Blida, gorges de la Chiffa. Corbière (1889), Jelenc (1955a): At-
(1951, 1967): Massif du Tessala, pic de Tafaraoui, 600 m, 650 m. las de Blida, source de Tala Yezid. Corbière (1904): Vallon d’El
Jelenc (1954b, 1967): Massif du Bou Zezga, gorges de Keddara, Biar près Alger. Corbière (1904), Jelenc (1955a): Jardin public
400 m. Jelenc (1955a): Atlas de Blida; Atlas de Blida, oued el Kebir, de Sidi-Bel-Abbès. Corbière (1905): Environs de Constantine,
300 m; Aumale; Biskra; Kabylie des Babor, cap Aokas; Kabyblie du Menidj; Environs de Constantine, Sidi M’Cid. Jelenc (1949,
Djurdjura, Aït Ali; Kabyblie du Djurdjura, Aït Ouaban; Kabylie 1955a, 1967): Monts de Tlemcen. Jelenc (1953, 1967): Aurès,
du Djurdjura, Akfadou; Kabylie du Djurdjura, Fort National; La flanc nord du djebel Chélia, 1800 m; Aurès, forêt de Sgag, 1650 m;
Réghaia; Monts de Tlemcen, Mazer; Ouarsenis, Téniet el Had. Bèlezma, djebel Refaa, 2100 m; Djurdjura, Aizer, en contre-bas
Jelenc (1955a, 1956, 1967): Sahel d’Alger, ravin du Frais-Vallon, du Chemin Lapie, 1750 m. Jelenc (1955a): Aïn Ouaban; Atlas
près d’El Biar, 300 m. Jelenc (1955a, 1967): Constantine. Jelenc de Blida, oued el Kebir, 200 m; Bône; Constantine; Kabylie des
(1955b, 1967): Haute-Plaine de Tlemcen. Jelenc (1966): Boghar, Babors, Sebt des Beni Slimane; Kabylie du Djurdjura, col de Ti-
djebel Krellala. Ros et al. (1990): Texenna’s wood, 700 m. Common. rourda, 1700 m; Littoral oranais; Nador de Médéa; Orléansville;
Philippeville. Jelenc (1956, 1967): Sahel d’Alger, dans le ravin
du Frais-Vallon. Jelenc (1966, 1967): Atlas Mitidjien, Hammam
Entosthodon obtusus (Hedw.) Lindb. Mélouane. Boukhatem et al. (2017): El-Kala National Park, Tonga
lake watershed. Common.
Records. — Bescherelle (1882), Jelenc (1955a): Blidah; Édough;
La Calle, bruyères; Oued-Amalou, route du Fort Génois; Oued-el-
-Aneb; Stora. Jelenc (1955a): Alger. Jelenc (1955a), Ahayoun et al. Comment
(2007): La Réghaia. Rare. Eucladium verticillatum var. angustifolium Jur. nom. illeg. was
reported from Algeria (Monts de Tlemcen, El Ourit) by Jelenc
(1949, 1967). Eucladium verticillatum var. angustifolium Lindb.
Entosthodon pulchellus (H.Philib.) Brugués was considered only a form of the type (Hodgetts et al. 2020)
and probably that is also the case.
Records. — Montagne (1838): Alger. Crundwell & Nyholm
(1974): Alger, ravin de Pierson; Constantine, Philippeville. Rare.

Eurhynchiastrum pulchellum
Entosthodon schimperi Brugués (Hedw.) Ignatov & Huttunen
Records. — Brotherus (1909), Jelenc (1955a): Algier. Very rare.
Eurhynchiastrum pulchellum (Hedw.) Ignatov & Huttunen var.
praecox (Hedw.) Ochyra & Zarnowiec

Ephemerum recurvifolium (Dicks.) Boulay Nomenclature follows Guerra (2016).

Records. — Montagne (1838): Alger. Bescherelle (1882), Jelenc

Records. — Bescherelle (1882): Alger. Bescherelle (1882), Jelenc (1955a): Tiaret. Lapie (1909): Kabylie du Djurjura. Jelenc (1955a):
(1955a): Birmandreïs; Philippeville. Rare. Kabylie du Djurdjura, Aizer. Rare.

Ephemerum serratum (Hedw.) Hampe

Eurhynchium striatum (Hedw.) Schimp.
Records. — Bescherelle (1882): La Calle; Mustapha. Bescherelle
(1882), Jelenc (1955a): Alger, à la Colonne Voirol. Rare. Records. — Bescherelle (1882), Jelenc (1955a): Kabylie du Djurd-
jura, Tizi Ouzou. Litschauer (1904): Qued-Kebir, 200 m. Jelenc
(1955a): Kabylie des Babor, cap Aokas. Rare.
Epipterygium tozeri (Grev.) Lindb.

Records. — Bescherelle (1882): Alger. Bescherelle (1882), Jelenc Fabronia leikipiae var. ahaggarensis Thér. & Trab.
(1955a): Beni-Salah; Bône; Bouzaréah; La Calle; Oued Amalou. Lapie
(1909): Kabylie du Djurjura. Jelenc (1951, 1967): Haute-Plaine de
Tlemcen, 650 m. Jelenc (1955a): Kabylie du Djurdjura, Akfadou; Records. — Thériot (1931, 1933), Jelenc (1955a): Atakor-n-
Kabylie du Djurdjura, Djemaà Saharidij; Kabylie du Djurdjura, Fort Ahaggar, in monte Amezzeroui, 2600 m. Very rare.
National; Kabylie du Djurdjura, Tifrit. Boukhatem et al. (2017):
El-Kala National Park, Tonga lake watershed. Frequent.
Comment
This taxon was described from Algeria by Thériot (1931).
Eucladium verticillatum (With.) Bruch & Schimp. According to Ros et al. (2013), its identity has not been
revised since. Fabronia leikipiae is mainly distributed in Africa
Records. — Montagne (1838), Bescherelle (1882), Jelenc (1955a): (O’Shea 2006) and recently has also been reported in Asia
Alger. Bescherelle (1882): La Chiffa; Médéah, fontaine Gros-Jean. (Yadav et al. 2019).

CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5) 75

 Ros R. M. et al.

Fabronia pusilla Raddi Fissidens curvatus Hornsch.

Records. — Bescherelle (1882): Gher-el Dabch. Bescherelle Records. — Corbière (1904), Jelenc (1955a): La Bouzaréah, près
(1882), Jelenc (1955a): Blidah, vieux oliviers; El-Harrouch, troncs Alger. Jelenc (1955a): Blida. Rare.
de lentisques; Milah; Mouzaïaville. Corbière (1905): Environs de
Constantine, route du Séminarie. Jelenc (1955a): Atlas de Blida;
Constantine; Kabylie du Djurdjura, Akfadou; Kabylie du Djurdjura, Fissidens dubius P.Beauv.
djebel Afroun. Jelenc (1955a), Ahayoun et al. (2007): Ouarsenis,
djebel Doui. Gauthier (1987): Assekrem, Hoggar. Boukhatem et al. Records. — Trabut (1914), Jelenc (1955a, 1956, 1967): Crêtes
(2017): El-Kala National Park, Tonga lake watershed. Frequent. du Djurdjura, Akfadou, 2000 m. Jelenc (1955a): Arba; Tabort
Amelel. Boukhatem et al. (2017): El-Kala National Park, Tonga
lake watershed. Rare.
Fissidens adianthoides Hedw.

Records. — Bescherelle (1882): Kabylie; Sud d‘Oran. Trabut Fissidens exilis Hedw.
(1887a): D’Oran à Mécheria. Jelenc (1955a): Arba; Djurdjura. Rare.
Records. — Corbière (1904), Jelenc (1955a): La Bouzaréah. Jelenc
Fissidens arnoldii R.Ruthe (1955a): Kabylie du Djurdjura, Aït Idjer. Rare.

Records. — Thériot (1931, 1933), Jelenc (1955a), Bruggeman- Fissidens fontanus (Bach. Pyl.) Steud.
Nannenga (1987): Hoggar, oued Haman, 2000 m. Very rare.
Records. — Bescherelle (1882): sine loco. Jelenc (1955a): La Ré-
Fissidens bryoides Hedw. ghaia; Laghouat. Jelenc (1955a), Ahayoun et al. (2007): Dayas de
l’Alma. Rare.
Records. — Desfontaines (1799): sine loco. Montagne (1838): Alger.
Pierrot (2000): Kabylie de Collo, berges de l’oued Melkach. Boukhatem
et al. (2017): El-Kala National Park, Tonga lake watershed. Rare. Fissidens grandifrons Brid.

Records. — Bescherelle (1882), Brongniart (1882), Jelenc (1955a):

Fissidens bryoides var. caespitans Schimp. Alger, petit Atlas de Blidah, cascades, 1200 m. Bescherelle (1882),
Jelenc (1955a): Fontaine de Gros-Jean, à Médéah, 1640 m; Gorges
de la Chiffa; Atlas de Blida, Oued-Kebir, 200 m. Corbière (1889),
Records. — Pierrot (2000): Kabylie de Collo, torrent de la source Jelenc (1955a): Atlas de Blida, source de Tala Yezid. Jelenc (1955a):
du Djebel Goufi. Very rare. Kabylie du Djurdjura, Afkadou; Mouzaïa. Rare.

Fissidens crassipes Wilson ex Bruch & Schimp.

Fissidens incurvus Starke ex Röhl.
Records. — Bescherelle (1882), Jelenc (1955a): Alger; Maison-Car-
rée, dans les ruisseaux; Stora. Jelenc (1955a): Bougie; El Biar; Kabylie Fissidens viridulus var. incurvus (Starke ex Röhl.) Waldh.
du Djurdjura, Aït Ouaban; La Réghaia. Jelenc (1951, 1955a, 1967):
Monts de Tlemcen, vallée de l’oued Khémis au village de Khémis; Nomenclature follows Hodgetts et al. (2020).
cascatelle près de la route, 800 m. Boukhatem et al. (2017): El-Kala
National Park, Tonga lake watershed. Rare. Records. — Bescherelle (1882), Jelenc (1955a): Alger, collines
ombragées; Bône. Corbière (1889): Blida. Corbière (1904):
Alger, colonne Voirol; Constantine, bois du djebel Ouasch;
La Bouzaréah, près Alger; Orléansville, bois de la Pépinière.
Fissidens crassipes subsp. warnstorfii Corbière (1905): Environs de Constantine, Les Pins, bords de
(M.Fleisch.) Brugg.-Nann. l’oued Bou Merzoug. Meylan (1931), Jelenc (1955a): Tlemcen.
Jelenc (1949, 1967): Vallée de l’Isser, source thermale de Sidi
Records. — Bescherelle (1882), Potier de La Varde (1922), Jelenc Abdelly, 375 m. Jelenc (1951, 1967): Haute-Plaine de Tlemcen,
(1955a): Bougie, ruisseaux. Potier de La Varde (1922), Jelenc (1955a): 700 m. Braun-Blanquet (1954): Forêt de Terni, près de Tlemcen,
La Réghaia. Jelenc (1949, 1967): Jardin d’Essai d’Alger. Jelenc (1949, 1000 m. Jelenc (1955a): Atlas de Blida; Constantine; Djurdjura;
1967), Bruggeman-Nannenga (1982): Monts et Haute-Plaine de Édough; El Affroun; La Bouzaréa; Orléansville. Jelenc (1956):
Tlemcen. Jelenc (1951): Monts de Tlemcen, cirque d’el Ourit. Fort des Arcades. Jelenc (1956, 1967): Massif du Bou Zegza,
Jelenc (1955a): La Bouzaréa. Jelenc (1955a), Bruggeman-Nannenga dans les Gorges de Keddara, 400 m; Plaine de la Mitidja, forêt
(1982): Kéddara. Rare. de la Règhaïa; Sahel d’Alger, environs d’El Biar; Sahel de Koléa,
au ravin des Voleurs. Common.

Fissidens crispus Mont. Fissidens ovatifolius R.Ruthe

Records. — Corbière (1904) Jelenc (1955a): Vallon d’El Biar Records. — Jelenc (1949, 1955a, 1967), Bonnot (1963): Monts
près Alger, 120 m. Jelenc (1954b, 1967): Sahel d’Alger, colline du de Tlemcen, forêt de Zariflet, 1100 m. Bonnot (1963): Oued
Hamma. Ros et al. (1990): Chiffa’s Gorge. Boukhatem et al. (2017): Yquem. Boukhatem et al. (2017): El-Kala National Park, Tonga
El-Kala National Park, Tonga lake watershed. Rare. lake watershed. Rare.

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 Revisiting the Bryophyte Checklist of Algeria

Fissidens pusillus (Wilson) Milde Kabylie du Djurdjura, Fort National, 1000 m; Monts de Tlemcen,
forêt de Zariffet; Monts de Tlemcen, forêt d‘Hafir; Sidi Merzoug.
Records. — Bescherelle (1882): Ben-Aknoun. Bescherelle (1882), Jelenc (1959): Kabylie de Collo, Baie de Tamamart, 50 m. Ros
Jelenc (1955a): Alger; Birkadem; Bou Khalfa; Kabylie; Stora. Gan- et al. (1990): Texenna’s wood, 700 m. Pierrot (2000): Kabylie de
doger (1883): sine loco. Corbière (1889), Jelenc (1955a): Blida. Collo. Boukhatem et al. (2017): El-Kala National Park, Tonga lake
Corbière (1905): Environs de Constantine, Le Hamma. Corbière watershed. Common.
(1905), Jelenc (1955a): Environs de Constantine, Sidi M’Cid, près les
Arcades romaines, pépinière du djebel Ouach. Jelenc (1951, 1967):
Haute-Plaine de Tlemcen, 700 m. Jelenc (1955a): Birmandreïs; El Fissidens viridulus (Sw. ex anon.) Wahlenb.
Biar; La Réghaia; Philippeville. Boukhatem et al. (2017): El-Kala
National Park, Tonga lake watershed. Frequent.
Fissidens exiguus Sull.
Fissidens subimmarginatus Philib.
Comment
According to Bruggeman-Nannenga (1978), F. subimmargi- Nomenclature follows Bruggeman-Nannenga (1978) and Hodgetts
natus Philib., as reported from the Massif de la Bouzaréa by et al. (2020).
Jelenc (1955b, 1967), should be regarded as a synonymous Records. — Braun-Blanquet & Maire (1924), Jelenc (1955a):
with F. exiguus. Later, Pursell (1997) considered F. exiguus Tlemcen. Jelenc (1949, 1951, 1955a, 1967): Haute plaine de
synonymous with F. bryoides var. pusillus (Wilson) Pursell, Tlemcen, berges du chabet bel Abbès, 775 m; Haute plaine de
and this was the criterion adopted by Ros et al. (1999) for the Tlemcen, 650-775 m. Jelenc (1954b, 1967): Sahel d’Alger, Qua-
checklis of Algeria. However, Ros et al. (2013) and Hodgetts tre-Chemins de Kouba. Jelenc (1955a): Maison-Carrée. Jelenc
(1955b, 1967): Massif de la Bouzaréa. Jelenc (1956, 1967): Sahel
et al. (2020) considered F. exiguus auct. eur. to be synomy- d’Alger, environs d’El Biar. Ros et al. (1990): Djebel Chenoua,
mous with F. viridulus, which is the criterion followed here. 5-40 m. Boukhatem et al. (2017): El-Kala National Park, Tonga
lake watershed. Rare.

Fissidens rivularis (Spruce) Schimp. Comment

Records. — Feldmann (1947), Jelenc (1955a): Kabylie de Collo, See comment under Fissidens pusillus.
vallée de l’oued Ramèche, 600-800 m. Very rare.

Flexitrichum flexicaule (Schwägr.) Ignatov & Fedosov

Fissidens rufulus Bruch & Schimp.
Ditrichum flexicaule (Schwägr.) Hampe
Records. — Corbière (1889), Jelenc (1955a), Ros & Cano (2008):
Blida. Corbière (1905), Jelenc (1955a): Environs de Constantine, Nomenclature follows Fedosov et al. (2016).
ravin du Rummel. Rare.
Records. — Bescherelle (1882): Djurjura; Kabylie. Bescherelle
(1882), Jelenc (1955a): Tlemcen. Jelenc (1953, 1955a, 1967):
Fissidens serrulatus Brid. Kabylie du Djurdjura, Aizer, falaises en contre-bas du Chemin
Lapie, 1750 m. Jelenc (1955a): Kabylie du Djurdjura, Aït Daoud;
Records. — Bescherelle (1882), Jelenc (1955a): Djebel Édough, Kabylie du Djurdjura, Tala Ghizan; Kef el Aïssa, Dépt. de Cons-
fond des ravins; Stora. Jelenc (1955a): Kabylie de Collo; Kabylie du tantine. Boukhatem et al. (2017): El-Kala National Park, Tonga
Djurdjura, Aït Daoud; Kabylie du Djurdjura, forêt d’Akfadou. Rare. lake watershed. Rare.

Fissidens taxifolius Hedw.

Fontinalis antipyretica Hedw. subsp. antipyretica
Fissidens taxifolius subsp. pallidicaulis (Mitt.) Mönk.
Fontinalis constantinica Cardot ex Jelenc nom. inval.
Nomenclture follows Hodgetts et al. (2020).
Nomenclature follows Welch (1960).
Records. — Bescherelle (1882): Petite Kabylie; Ras-Aokas. Besche-
relle (1882), Jelenc (1955a): Bône; Djebel Édough, près la Fontaine Records. — Bescherelle (1882), Jelenc (1955a): Alger, Tixeraïn.
du Prieur. Corbière (1904), Jelenc (1955a): La Bouzaréah, près Alger. Bescherelle (1882), Welch (1960): Kabylie. Julien (1894): Région
Corbière (1905), Jelenc (1955a): Environs de Constantine, djebel de Constantine. Lapie (1909): Kabylie du Djurjura. Trabut (1914),
Ouach, 900 m. Camus (1906): Oran. Meylan (1931): Tlemcen. Jelenc (1955a), Ahayoun et al. (2007): Ruisseaux de la forêt d’Akfadou.
Feldmann (1947), Jelenc (1955a): Kabylie de Collo, vallée de l‘oued Jelenc (1955a): Babor; Kabylie du Djurdjura, Taourirt Ighil; Kabylie
Bin el Ouidane, près de Taoulel, 600 m. Dubuis & Faurel (1945), du Djurdjura, Tirourda; Kabylie du Djurdjura, Yakouren; Monts de
Jelenc (1953, 1955a): Djurdjura, Aizer, flanc nord du djebel Tach- Constantine, djebel Ouach; Oued Réghaia. Jelenc (1955a), Welch
gagalt, 2000 m. Dubuis & Faurel (1945), Jelenc (1953, 1955a, (1960): La Calle. Jelenc (1955a), Ahayoun et al. (2007): Kabylie
1967): Kabylie du Djurdjura, Plateau de l‘Aizer, 1900-2000 m. du Djurdjura, Tala Ghizan. Jelenc (1955a), Ros & Cano (2008):
Braun-Blanquet (1954): Forêt de Terni, près de Tlemcen, 1000 m. Kabylie du Djurdjura, Tamesguida. Welch (1960): Tiaret. Welch
Jelenc (1954b, 1967): Massif du Bou Zegza, gorges de Keddara, (1960), Jelenc (1967): Constantine. Boukhatem et al. (2017): El-Kala
400 m. Jelenc (1955a), Pierrot (2000): Alger. Jelenc (1955a): Blida; National Park, Tonga lake watershed. Mazari & Boulaacheb (2023):
Kabylie des Babors, cap Aokas; Kabylie du Djurdjura, Akfadou; Setif, Megriss Mountain. Frequent.

CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5) 77

 Ros R. M. et al.

Fontinalis hypnoides var. duriaei (Schimp.) Kindb. Forêt de Terni, près de Tlemcen. Léredde (1954): Tassili N’Aijer.
Jelenc (1951, 1967): Djebel Borouse; Sahel d’Arzeu, plateau du djebel
Records. — Schimper (1876), Bescherelle (1882), Jelenc (1955a), Orouse, 630 m; Sahel d’Oran, plateau des Ghamras, 400 m. Jelenc
Welch (1960): Prope La Calle ad ripas lacus el Hout Algeriae in (1954b, 1967): Massif du Bou Zezga, gorges de Keddara, 400 m;
Alni. Bescherelle (1882), Jelenc (1955a), Welch (1960): Alger, Bas- Monts de Tlemcen, Massif de Ghar Rouban, l’oued Zaouia; Chaîne
sin de la fontaine du Café maure de Kaddous. Jelenc (1951, 1967): des Zahrez, au nord du Zahrez Chergui, djebel Touilila, 1000 m,
Monts de Tlemcen, forêt de Zariflet, 1000-1100 m, cascades de l’oued M’Kamza. Jelenc (1955a): Atlas de Blida, oued el Kebir, 200 m;
l’oued Ouadallah. Jelenc (1955a): Kabylie du Djurdjura, Akfadou; Djemila; Hauts-plateaux oranais; Kabylie du Djurdjura, djebel Afroun;
Tiaret, oued Mina; Tixerain. Jelenc (1955a), Welch (1960): Oued Kabylie du Djurdjura, Fort National; Marengo; Miliana; Ouarsenis;
Réghaia, barrage; Oued Saoula. Jelenc (1955a), Ahayoun et al. Tipaza, Tombeau de la Chrétienne; Tlemcen. Boukhatem et al. (2017):
(2007): Birtouta. Rare. El-Kala National Park, Tonga lake watershed. Mazari & Boulaacheb
(2023): Setif, Megriss Mountain. Common.

Fontinalis squamosa Hedw. Comment

Two invalid names were used in Algeria by Jelenc (1955a)
Records. — Bescherelle (1882), Jelenc (1955a): sine loco. Very rare. whose identity is unknown: F. hygrometrica var. brevipes Jelenc
nom. nud. (reported from Alger and Blida) and F. hygrometrica
Comment var. macrocarpa Jelenc nom. nud. from Hauts-plateaux oranais.
The only report of the species in Algeria is based on a single
specimen collected by Durieu de Maisonneuve.
Funaria microstoma Bruch ex Schimp.

Funaria altissima Dixon Records. — Jelenc (1955b, 1967): Massif de la Bouzaréa. Boukhatem
et al. (2017): El-Kala National Park, Tonga lake watershed. Rare.
Records. — Dixon (1931), Ros et al. (2023): Taharauet, south-
ern part of Hoggar Mts., Central Sahara, 5900 ft. Jelenc (1955a):
Hoggar méridional, Fort Motylinski. Very rare. Funariella curviseta (Schwägr.) Sérgio

Records. — Montagne (1838), Bescherelle (1882), Jelenc (1955a):

Comment Alger. Bescherelle (1882): Oran. Bescherelle (1882), Jelenc (1955a):
Blidah; La Calle; Philippeville. Trabut (1887a): D‘Oran à Méche-
According to Ros et al. (2023), the locality in Jelenc (1955a) ria. Corbière (1904): Alger, talus humide du chemin de Telemly,
is with high probability the same as in the protologue. Cur- 80 m. Camus (1906): environs de Tlemcen. Jelenc (1954b, 1967):
rently, it is a species endemic to Algeria. Massif du Bou Zezga, gorges de Keddara, 400 m. Jelenc (1955a):
Baba Ali; Birmandreïs; Kabylie du Djurdjura, Aït Idjer; Kabylie du
Djurdjura, Tala Ghizan; Nador de Médéa. Jelenc (1955a), Ahayoun
et al. (2007): Bouzaréah. Boukhatem et al. (2017): El-Kala National
*Funaria flavicans Michx. Park, Tonga lake watershed. Frequent.
Records. — Montagne (1838): Alger.
Grimmia anodon Bruch & Schimp.
Comment
Doubtfully present in Algeria. The single report of Montagne Records. — Bescherelle (1882): Blidah, Petit Atlas, Sommet de
(1838) was not included in the checklist of Jelenc (1955a) l’Atlas. Jelenc (1953, 1967): Massif de l’Aurés, sommet du djebel
Lazreg, 1900 m. Jelenc (1955a): Atlas de Blida; Bou Saâda; Monts
and Ros et al. (1999, 2013). The presence in Algeria of this de Ksour, djebel Mzi. Boukhatem et al. (2017): El-Kala National
species which is mainly distributed in North America and also Park, Tonga lake watershed. Rare.
present in Japan (Miller & Miller 2007; Kiguchi & Sawada
2020) is highly doubtful.
Grimmia capillata De Not.

Funaria hygrometrica Hedw. Records. — Coppey (1911), Trabut (1927), Jelenc (1950, 1955a):
Ravin de Ben Zireg près Beni-Ounif, dans l’extrème-Sud Oranais.
Records. — Desfontaines (1799), Gandoger (1883), Ahayoun et al. Jelenc (1950, 1951, 1967): Environs d’Hadjerat el Mguil, 950 m;
(2007): sine loco. Montagne (1838), Bescherelle (1882), Jelenc (1955a): Hadjerat el Mguil. Jelenc (1950, 1955a): Entre Ghardaia et El Goléa.
Alger. Bescherelle (1882): Barbarie; Djurjura; Sud d‘Oran. Bescherelle Jelenc (1951: Atlas saharien (région de Figuig), djebel Grouz au cirque
(1882), Corbière (1889): Blidah. Bescherelle (1882), Jelenc (1955a): de Djahfat, 1400 m. Jelenc (1955a): Mzab. Frahm (1978): Laghouat.
Bône; Édough; Kabylie du Djurdjura, Tizi Ouzou; Sidi-Ferruch. Trabut Greven (1995): Bani Oussif, Vivgargel; Biscra, Col de Sfa. Boukhatem
(1887a): D’Oran à Mécheria, Chott. Julien (1894): Région de Con- et al. (2017): El-Kala National Park, Tonga lake watershed. Rare.
stantine. Corbière (1904), Jelenc (1955a): Cherchell. Corbière (1905),
Jelenc (1955a): Environs de Constantine, La Corniche; Environs de
Constantine, Moulin Laire; Environs de Constantine, Pépinière du Grimmia crinita Brid.
djebel Ouach. Camus (1906), Jelenc (1955a): Oran. Lapie (1909):
Kabylie du Djurjura. Thériot (1931, 1933), Jelenc (1955a), Ros & Records. — Bescherelle (1882): Chardina; Djebel Gharribou; El-Outaïa;
Cano (2008): Ahaggar, Ideles, 1450-1500 m. Braun-Blanquet (1954): Mont de Sel. Bescherelle (1882), Jelenc (1955a): Monts de Constantine;

78 CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5)

 Revisiting the Bryophyte Checklist of Algeria

Oran, colonie agricole du Saint-Denis-du-Sig; Plateau de Mansourah; Mascara. Jelenc (1955a): Kabyblie du Djurdjura, Aït Idjer; La Calle;
Tiaret. Bescherelle (1882), Frahm (1978): Biskra. Trabut (1887a): Lac el Hout; Palestro. Rare.
D’Oran à Mécheria. Julien (1894): Région de Constantine. Jelenc
(1955a): El Kantara; Tlemcen. Greven (1995): Laghouat. Boukhatem
et al. (2017): El-Kala National Park, Tonga lake watershed. Frequent. Comment
After the recognition of G. dissimulata E.Maier (Maier 2002),
Comment it is highly probable that some reports of G. meridionalis actu-
Grimmia crinita var. elongata Bruch & Schimp. was reported ally pertain to the former species.
by Camus (1906) from the environs de Tlemcen and by Jelenc
(1955a) from Aumale and Biskra. Although Index Muscorum
(Wijk et al. 1962) stated that this is a synonym of Grimmia Grimmia montana Bruch & Schimp.
capillata De Not., Muñoz & Pando (2000) considered the
identity of the name uncertain. Records. — Jelenc (1955a): El Outaya. Very rare.
Grimmia crinita Brid. f. compacta Besch. nom. nud. was
reported by Jelenc (1955a) from Outaia. Muñoz & Pando
(2000) could not locate the original material and its identity Grimmia orbicularis Bruch ex Wilson
has not been revised (Ros et al. 2013).
Records. — Bescherelle (1882): Aïn Timendert; Oran, hauts
plateaux, entre Chellala Dahrania et Arba Tahtani; Sahara algérien.
Bescherelle (1882), Meylan (1931): Constantine, sommet du
Grimmia decipiens (Schultz) Lindb. Mansourah. Bescherelle (1882), Jelenc (1955a): Aurès; Bou-Sâada;
Djelfa; Guelt es Stel. Bescherelle (1882), Jelenc (1955a), Frahm
Records. — Corbière (1904), Jelenc (1955a): Médéa. Jelenc (1953, (1978): Laghouat. Trabut (1887a): D‘Oran à Mécheria. Corbière
1967): Massif de Bélézma, cédraies de l’oued Chaba, 1400 m; Massif (1889): Blida. Corbière (1904), Jelenc (1955a): Batna, rochers
de l’Aurès, forêt de Sgag, 1700 m. Jelenc (1955a): Atlas de Blida, calcaires sur le flanc sud du djebel Tougourth, 1600 m; Médéa; Tia-
oued el Kebir, 300 m; Kabylie des Babor, cap Aokas; Kabylie du ret, région des Hauts-Plateaux, plaine du Douar Torrich, 1000 m.
Djurdjura, Fort National; Kabylie du Djurdjura, Michelet, 1000- Corbière (1905): Environs de Constantine, Aïn Yagout; Environs
1200 m. Rare. de Constantine, blocs calcaires au bord de l‘oued M‘Gharoud;
Environs de Constantine, Les Pins; Environs de Constantine,
Sidi M‘Cid. Corbière (1905), Jelenc (1955a), Muñoz & Pando
(2000): Constantine, Chettabah, près l‘usine Micout. Coppey
Grimmia funalis (Schwägr.) Bruch & Schimp. (1911), Jelenc (1955a): Beni-Ounif, Sahara algérien. Jelenc (1951,
1967): Monts de Tlemcen, djebel el Hassi, 1100 m. Jelenc (1953,
Records. — Bescherelle (1882), Jelenc (1955a): Aurès. Jelenc 1967): Massif de l‘Aurès, flanc nord du djebel Pharaoun, 1800 m.
(1955a): Atlas de Blida, 1400 m. Mazari & Boulaacheb (2023): Braun-Blanquet (1954), Jelenc (1955a): Tlemcen, 1000 m. Jelenc
Setif, Megriss Mountain. Rare. (1954b, 1967): Chaîne des Zahrez, au Nord du Zahrez Chergui,
crête du Draa Ferchet-el-Arbi. Jelenc (1955a): Atlas de Blida, oued
el Kebir; Biban; Boghar; Djebel Amour, Aïn Timandert; Djebel
Grimmia hartmannii Schimp. Dira; El Guerrah; El Kantara; Gorges de la Chiffa, 300 m; Oran,
Monts de Tlemcen; Ouarsenis; Reibell; Saïda; Sud constantinois,
Records. — Jelenc (1955a): Souk-Ahras. Very rare. Biskra; Tebessa; Téniet Zebboudj. Jelenc (1956, 1967): Massif
du Bou Zegza, dans les gorges de Keddara, 400 m. Jelenc (1966,
1967): Zahrez Chergui, oued Mlilia au djebel Touilila, 1000 m.
Bizot (1973): Fontaine Chaude à 30 km NE de Batna; Ruines de
Grimmia laevigata (Brid.) Brid. Timgad. Boukhatem et al. (2017): El-Kala National Park, Ton-
ga lake watershed. Mazari & Boulaacheb (2023): Setif, Megriss
Records. — Bescherelle (1882): Bab-Azoun; Vallée de l’oued Kou- Mountain. Very common.
bba. Bescherelle (1882), Jelenc (1955a): Alger; Édough; Philippeville;
Pic Belloua; Tizi-Ouzou; Vallon de Birkadem. Gandoger (1883): Comment
sine loco. Corbière (1905), Jelenc (1955a): Environs de Constantine,
djebel Ouach. Jelenc (1953, 1967): Massif du Bélezma, crêtes de Grimmia orbicularis var. longipila Husn. was reported by
Bordjem, 2200 m. Jelenc (1955a): Akfadou; Atlas de Blida, oued el Jelenc (1955a) from several localities (Aumale; Coll d’Aïn
Kebir, 300 m; Bône; Kabylie du Djurdjura, Fort National, 1000 m; Kebira; El Aricha; El Kantara; Tiaret) but according to Ros
Médéa. Frequent. et al. (2013), its identity has not been revised since. Muñoz &
Pando (2000) could not locate the original material.
Grimmia lisae De Not.

Records. — Montagne (1856): Circa lacum El Hout. Jelenc Grimmia ovalis (Hedw.) Lindb.
(1955a): Constantine. Jelenc (1955b, 1967): Bône; La Calle. Rare.
Records. — Bescherelle (1882): Petit Atlas. Bescherelle (1882),
Jelenc (1955a): Djebel Ksel, près de Géryville; Kabylie du Djurdju-
Grimmia meridionalis (Müll. Hall.) E.Maier ra, Fort National, 1000 m. Corbière (1889), Jelenc (1955a): Atlas
de Blida, Beni-Salah, région des Cèdres. Lapie (1909): Kabylie
Records. — Bescherelle (1882), Corbière (1904), Jelenc (1955a): du Djurjura. Jelenc (1953, 1955a, 1967): Massif de l’Aurès, flanc
Bône; Djebel Édough. Bescherelle (1882), Jelenc (1955a): Alger; nord du Chélia, 1700 m; Massif de l’Ouarsenis, cédraie de Téniet

CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5) 79

 Ros R. M. et al.

el Had, 1500 m. Jelenc (1953, 1967): Aurès, Ras-el-Ma, flanc nord Grimmia trichophylla Grev.
du djebel Pharaoun, 1800 m; Kabylie du Djurdura, Aizer, Aux
Beni Kouffi, 1600 m; Massif de l’Ouarsenis, Kef Siga, 1700 m; Records. — Bescherelle (1882), Jelenc (1955a): Édough, rochers
Massif du Bèlezma, crêtes de Bordjem, 2200 m. Jelenc (1955a): des caroubiers. Trabut (1887a): D’Oran à Mécheria. Jelenc (1955a):
Akfadou; Bône; Col de Tirourda, 1700 m; Djebel Amour; Dra Boghar; Kabylie du Djurdjura, Akfadou; Kabylie du Djurdjura, Fort
Inguel; Édough; Hauts-plateaux oranais; Orléansville; Tiaret. Jelenc National; Mascara; Oran; Palestro; Portes de Fer. Rare.
(1955b): Camp-du Maréchal au marabout de Sidi Ali bou Nab,
885 m; Kabylie du Djurdjura, N d’Aizer, 1500 m. Jelenc (1967):
Camp du Maréchal, 885 m. Bizot (1973): Djebel Belezma à 15 km
à l’ouest de Batna. Mazari & Boulaacheb (2023): Setif, Megriss Gymnostomum aeruginosum Sm.
Mountain. Common.
Records. — Jelenc (1955a): Kabylie du Djurdjura, gorges de Tifrit,
1340 m. Very rare.
Grimmia pulvinata (Hedw.) Sm.

R ecords . — Montagne (1838), Bescherelle (1882), Jelenc Gymnostomum calcareum Nees & Hornsch.
(1955a): Alger. Bescherelle (1882): Chatabch; Col d’Aïn Kebi-
ra; Petit Atlas; Tiaret; Tlemcen. Bescherelle (1882), Brongniart Records. — Corbière (1904), Camus (1906), Jelenc (1955a):
(1882): Blidah. Bescherelle (1882), Corbière (1904), Jelenc Tlemcen. Corbière (1904), Jelenc (1955a): Alger; Blida; Tiaret,
(1955a): Médéah. Bescherelle (1882), Corbière (1905): Con- Hauts-Plateaux. Corbière (1905): Environs de Constantine, Les
stantine. Bescherelle (1882), Jelenc (1955a): Aurès; Bouzaréah; Pins; Environs de Constantine, route de la Corniche. Corbière
Philippeville; Tizi-Ouzou. Gandoger (1883): sine loco. Corbière (1905), Whitehouse & Crundwell (1992): Environs de Con-
(1889): Blida, oued Beni-Aza; Blida, près du marabout Sidi-Mo- stantine, Sidi M’Cid. Jelenc (1951, 1967): Massif du Tessala,
hamed-er-Rarbi. Cardot (1904), Maire (1916), Jelenc (1955a): pic de Tafaraoui, 600 m. Jelenc (1954b, 1967): Sahel d’Alger,
Province d’Oran, Djebel Aïssa, 1800 m, 2250 m. Corbière (1904), colline du Hamma. Jelenc (1954b, 1955a, 1967): Kabylie du
Jelenc (1955a): Sétif, 1100 m. Corbière (1905), Jelenc (1955a): Djurdjura, Monts de Saida, cascades de Tifrit. Jelenc (1955a):
Environs de Constantine, Les Pins; Environs de Constantine, Sidi Alger, Frais Vallon; Arba; Atlas de Blida, oued el Kebir, 200 m;
M’Cid. Camus (1906): Oran. Braun-Blanquet & Maire (1924): Constantine; Djebel Chenoua; Kabylie du Djurdjura, Aït Idjer;
Tlemcen, 800-900 m. Jelenc (1951): Massif du Tessala, pic de Khodjaberry; Mascara; Médéa; Monts de Tlemcen, Mazer; Zac-
Tafaraoui, 650 m. Jelenc (1951, 1967): Monts de Tlemcen, djebel car. Jelenc (1956, 1967): Sahel d’Alger, dans environs d’El Biar.
el Beniane, 1100 m. Jelenc (1953): Aurès, flanc nord du Chélia, Ros et al. (1990): Chiffa’s Gorge. Whitehouse & Crundwell
1700 m; Flanc nord du djebel Pharaoun, 1800 m; Sommet du (1992): Cascades of El Ourit, Tlemcen; Constantine, chemin
djebel Lazreg, 1900 m. Jelenc (1953, 1967): Maadid, maison des Touristes. Boukhatem et al. (2017): El-Kala National Park,
forestiére de Mahali, 1300 m; Massif de Guethiane, Kef Tachrirt, Tonga lake watershed. Mazari & Boulaacheb (2023): Setif, Me-
1850 m. Jelenc (1955a): Aïn el Hadjar; Akfadou; Atlas de Blida, griss Mountain. Common.
oued el Kebir, 300 m; Azéba; Batna; Col de Tirourda, 1700 m;
Djebel Chenoua; Gorges de la Chiffa; Guelt es Stel; Kabylie du
Djurdjura, Afensou; Kabylie du Djurdjura, Fort National, 1000- Comment
1200 m; Lambèse; Monts de Ksour, djebel Mzi; Ouarsenis, Téniet
el Had Tablabalt. Jelenc (1956, 1967): Atlas Mitidjien, pic Mou-
Gymnostomum calcareum var. gracillimum Besch. nom. nud.
zaïa; Massif du Bou Zegza, dans les gorges de Keddara, 400 m. was reported by Bescherelle (1882) and Jelenc (1955a) from
Jelenc (1967): Massif de l’Aurès, 1700-1900 m. Jelenc (1967), Alger. According to Ros et al. (2013), the identity of this name
Ahayoun et al. (2007): Aumale. Bizot (1973): Djebel Belezma à seems not to have been revised since. It is not a synonym of
15 km à l’ouest de Batna. Mazari & Boulaacheb (2023): Setif, Hymenostylium gracillimum (Nees & Hornsch.) Köckinger &
Megriss Mountain. Very common.
Jan Kučera.

Comment
Grimmia pulvinata var. longipila Schimp. was reported by Gymnostomum viridulum Brid.
Jelenc (1955a) from Monts de Constantine and by Jelenc
(1967) from Massif du Tessala, pic de Tafaraoui, 650 m. Records. — Bescherelle (1882): Birkadem; Birmandreïs, rochers
According to Ros et al. (2013), its identity has not been ombragés; Mustapha. Bescherelle (1882), Jelenc (1955a): Alger;
revised since. Muñoz & Pando (2000) could not locate the Constantine, Chute du Rummel; Mansourah; Mascara. Trabut
original material. (1887a), Jelenc (1955a): D’Oran à Mécheria, Tell. Jelenc (1955a),
Ros et al. (1990): Djebel Chenoua. Boukhatem et al. (2017): El-
Kala National Park, Tonga lake watershed. Rare.

Grimmia teretinervis Limpr.

Records. — Jelenc (1955a): Kabylie du Djurdjura, Fort National, Gyroweisia reflexa (Brid.) Schimp.
1000 m. Very rare.
Records. — Montagne (1838), Jelenc (1955a): Alger. Bescherelle
(1882): Alger, mur humide de la Salpétrière. Bescherelle (1882),
Grimmia tergestina Tomm. ex Bruch & Schimp. Corbière (1905), Jelenc (1955a): Environs de Constantine, au bord
du oued bou Merzoug. Bescherelle (1882), Jelenc (1955a): Blidah.
Records. — Corbière (1905), Jelenc (1955a): Environs de Con- Corbière (1905): Environs de Constantine, Telemly près Alger.
stantine, Les Pins. Very rare. Jelenc (1955a): Azéba. Rare.

80 CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5)

 Revisiting the Bryophyte Checklist of Algeria

Habrodon perpusillus (De Not.) Lindb. Homalia trichomanoides (Hedw.) Schimp.

Records. — Bescherelle (1882): Blidah; Kabylie. Bescherelle Records. — Jelenc (1955a) sine loco. Jelenc (1955b, 1967) Con-
(1882), Jelenc (1955a): Kabylie du Djurdjura, près de pic Belloa, stantine. Boukhatem et al. (2017): El-Kala National Park, Tonga
1550 m; Kabylie du Djurdjura, Tizi-Ouzou. Gandoger (1883): sine lake watershed. Rare.
loco. Corbière (1889), Jelenc (1955a): Atlas de Blida, forêt des Beni
Salah. Jelenc (1951, 1967): Monts de Tlemcen, forêt de Zariflet,
près de l’Aïn Defla. Jelenc (1955a): Alger; Kabyblie du Djurdjura, Homalothecium aureum (Spruce) H.Rob.
Aït Ali; Kéddara. Jelenc (1956, 1967): Atlas Mitidjien, près du Col
des Deux-Bassins. Jelenc (1966, 1967): Monts de Constantine au Records. — Durieu de Maisonneuve (1849), Gandoger (1883),
djebel Ouach, 1000 m. Boukhatem et al. 2017): El-Kala National Trabut (1888): sine loco. Bescherelle (1882): Djebel Beni-Salah;
Park, Tonga lake watershed. Frequent. Djurjura; Kabylie. Bescherelle (1882), Litschauer (1904), Jelenc
(1955a): Fort National und Michelet, 1000-1200 m. Bescherelle
(1882), Jelenc (1955a): Alger, près de la Maison-Carrée; Bouïroun;
*Hamatocaulis vernicosus (Mitt.) Hedenäs Mascara; Tiaret. Brongniart (1882): Blidah. Trabut (1887a): D’Oran
à Mécheria, Tell. Corbière (1904): Tiaret, Hauts-Plateaux, Douar
Records. — Jelenc (1955a): Alger; Gorges de la Chiffa. Torrich. Corbière (1904), Jelenc (1955a): Médéa, bois de Tibarrine;
Orléansville, bois de la Pépinière. Litschauer (1904), Jelenc (1955a):
Chiffaschlucht bei Blidah, 200 m. Corbière (1905), Jelenc (1955a):
Comment Environs de Constantine, Chettabah; Environs de Constantine,
Doubtfully present in Algeria. The presence of this Cir- djebel Ouach; Environs de Constantine, Les Pins; Environs de
cumpolar Boreal-montane species (Blockeel et al. 2014b) in Constantine, route de Sétif; Environs de Constantine, route du
Séminarie. Lapie (1909): Kabylie du Djurjura. Jelenc (1955a): Alger;
Algeria is unlikely in the reported locality. Given the chal- Atlas de Blida; Monts de Tlemcen, djebel Tenouchfi; Tala Ghizan.
lenges in locating these collections, this species could only Hofmann (1998): Algier, Ojebelonach foret de l’cOru. Boukhatem
be retained in the Algerian catalogue upon the discovery of et al. (2017): El-Kala National Park, Tonga lake watershed. Mazari &
new specimens. Boulaacheb (2023): Setif, Megriss Mountain. Frequent.

Hedwigia ciliata (Hedw.) P.Beauv. Homalothecium lutescens (Hedw.) H.Rob.

Records. — Bescherelle (1882), Jelenc (1955a): Bône; Djebel Records. — Litschauer (1904): Chiffaschlucht bei Blidah, 200 m;
Mouzaïa; Djurdjura; Kabylie du Djurdjura, Tizi Ouzou; Massif de la Kabylie du Djurdjura, Tirourda, 1700 m. Jelenc (1949, 1967):
Mahouna, Serdj el Aouda. Jelenc (1955a): Atlas de Blida; Djidjelli; Haute-Plaine de Sétif, Ain Trich, 1100 m. Jelenc (1955a): Atlas de
Édough; Kabylie du Djurdjura, Akfadou. Rare. Blida, oued el Kebir, 200 m; Kabylie du Djurdjura, Aizer; Médéa;
Monts de Tlemcen, djebel Tenouchfi. Rare.

Hedwigia emodica Hampe ex Müll. Hal.

Homalothecium lutescens var. fallax H.Philib. ex Schimp.
Hedwigia ciliata var. leucophaea Bruch & Schimp.
Nomenclature follows Ignatova et al. (2016). Records. — Corbière (1904): Médéa. Very rare.

Records. — Jelenc (1955a): Édough. Very rare.

Comment
This variety was not included in Ros et al. (1999) because
it was not accepted as different from var. lutescens by Jelenc
Herzogiella seligeri (Brid.) Z.Iwats. (1955a) and was overlooked. Nor did Ros et al. (2013) con-
sider this taxon for Algeria.
Records. — Bescherelle (1882): sine loco. Jelenc (1955a): Atlas
de Blida. Very rare.

Homalothecium philippeanum (Spruce) Schimp.

Homalia lusitanica Schimp.
Records. — Bescherelle (1882): Djurjura, près Bordj Boghni;
Records. — Bescherelle (1882), Corbière (1904), Jelenc (1955a), Kabylie; Taguemount-Azouz. Bescherelle (1882), Jelenc (1955a):
Ahayoun et al. (2007): Gorges de la Chiffa, ruisseau des Singes. Aurès; Blidah, petit Atlas, rochers; Bouzaréah. Bescherelle (1882),
Bescherelle (1882), Jelenc (1955a): Djebel Édough, fond des Jelenc (1955a), Hofmann (1998): Aurès. Corbière (1904, 1905):
ravins, au nord. Trabut (1888): sine loco. Lapie (1909): Kabylie Constantine, djebel Ouasch, 1100 m, et rochers calcaires sur la route
du Djurjura. Battandier et al. (1914), Jelenc (1955a): Forêt des de la Corniche. Jelenc (1953, 1955a, 1967): Massif de l’Ouarsenis,
Aït-Ouaban, groute l’Ifri-bou-Anou. Feldmann (1947), Jelenc cédraie de Téniet el Had, 1500 m. Jelenc (1953, 1967): Bèlezma,
(1955a, 1959): Kabylie de Collo, Baie de Tamanart, moins de cédraies de l’oued Chaba, 1400 m; Massif de l’Aurès, flanc nord
50 m. Jelenc (1954b, 1967): Sahel de Koléa, au ravin des Voleurs. du djebel Pharaoun, 1800 m; Massif de l’Aurès, flanc sud du djebel
Jelenc (1955a): Arba; Atlas de Blida; Djebel Chenoua; Kabylie Chélia, 1700 m; Massif de l’Aurès, forêt de Sgag, 1700 m; Monts
du Djurdjura, forêt des Aït Ouaban; Kabylie du Djurdjura, Ifir du Hodna, Massif de Guethiane au Kef Tachrit, 1850 m. Jelenc
Smeden. Boukhatem et al. (2017): El-Kala National Park, Tonga (1955a): Boghar; Constantine. Jelenc (1966, 1967): Kabylie du
lake watershed. Frequent. Djurdjura à Aït Ouabane. Bizot (1973): Djebel Belezma à 15 km à

CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5) 81

 Ros R. M. et al.

l’ouest de Batna. Boukhatem et al. (2017): El-Kala National Park, Hydrogonium bolleanum (Müll. Hal.) A.Jaeger
Tonga lake watershed. Common.
Barbula bolleana (Müll. Hal.) Broth.
Comment Barbula ehrenbergii var. algeriae (Müll. Hal. ex Geh.) Latzel
Homalothecium philippeanum var. kabylica Trab. ex Jelenc
nom. inval. descr. gall. was reported by Jelenc (1955a) from Didymodon mediterraneum var. algeriae (taxon not formally published
but name used by Corbière 1905).
Kabylie du Djurdjura, Taguemmount-Azouz, but according
to Ros et al. (2013), its identity has not been revised since. Hydrogonium ehrenbergii var. algeriae (Müll. Hal. ex Geh.) Podp.
Hyophila ehrenbergii var. algeriae (Müll.Hal. ex Geh.) Pavletic
Homalothecium sericeum (Hedw.) Schimp. Trichostomum mediterraneum var. algeriae Müll. Hal. ex Geh.

R ecords . — Desfontaines (1799), Gandoger (1883), Tra- Nomenclature follows Kučera et al. (2013) and Ros et al. (2023).
but (1888): sine loco. Montagne (1838), Jelenc (1955a): Al- Records. — Philibert (1880a), Jelenc (1955a): Gorges de la Chiffa,
ger. Bescherelle (1882): Alger, Atlas; Constantine; Djurjura. près de Blidah. Corbière (1904): Tlemcen, cascatelle sur la route des
Bescherelle (1882), Jelenc (1953, 1955a, 1967): Kabylie des cascades. Jelenc (1951, 1967): Haute plaine de Tlemcen, cascades
Babor, crête du djebel Tababor, 1950 m. Bescherelle (1882), de l’oued Bou Ennag à la traversée des collines des Beni Mester.
Jelenc (1955a): Édough; Kabylie du Djurdjura, Tizi Ouzou; La Jelenc (1954b, 1967): Monts de Saida, cascades de Tifrit. Jelenc
Calle; Philippeville. Corbière (1889): Blida. Corbière (1904), (1955a): Kabylie du Djurdjura, Palestro; Mazer; Monts de Tlemcen,
Jelenc (1955a): Gorges de la Chiffa, ruisseau des Singes; Mé- Tlemcen. Boukhatem et al. (2017): El-Kala National Park, Tonga
déa. Litschauer (1904): Qued-Kebir, 200 m. Litschauer (1904), lake watershed. Rare.
Jelenc (1955a): Desgleichen bei Fort National, 1000 m; Kabylie
du Djurdjura, col de Tirourda, 1700 m. Corbière (1905), Jelenc
(1955a): Environs de Constantine, Sidi M’Cid. Camus (1906), Comment
Jelenc (1955a): Oran. Lapie (1909): Kabylie du Djurjura. Braun- Barbula ehrenbergii var. immersa Trab. ex Jelenc nom. inval.
Blanquet & Maire (1924): Tlemcen. Maire & Sevenet (1928),
Jelenc (1955a): Tombeau de la Chrétienne, près de Bérard. Jelenc was proposed by Jelenc (1955a) from Sebdou. According
(1949, 1955a, 1967): Monts de Tlemcen, Sidi Djilali; Monts to Ros et al. (2013), the identity of this name has not been
de Tlemcen, Tlemcen. Jelenc (1951, 1967): Massif du Tessala, revised since its description.
pic Tessala, 1000 m. Jelenc (1953, 1955a, 1967): Kabylie du Barbula trabutii Jelenc nom. inval. descr. gall. was proposed
Djurdjura, massif de l’Aizer, fissures des falaises en contre-bas du by Jelenc (1955a) for the specimens collected and identified by
Chemin Lapie, 1750 m; Massif de l’Aurès, flanc nord du djebel
Chélia, 1700 m. Jelenc (1953, 1967): Atlas de Blida, ravin Bleu Corbière as Didymodon trabutii Corb. ex Jelenc nom. inval.,
de Chréa; Massif du Bélézma, cédraies de l’oued Chaba, 1400 m. cited as a synonym, that was sent to Trabut and remained
Jelenc (1955a): Atlas de Blida, oued el Kebir; Bou Saâda, djebel unpublished. Jelenc (1955a) indicated that this name could
Fernane, 1600 m; Djebel Chenoua; Djebel Dira; Djebel Doui; be just a local variation of Barbula ehrenbergii (synonym:
Kabylie des Babor, Djidjelli; Kabylie des Babor, Dra el Arba;
Kabylie du Djurdjura, Aït Khalfoun; Kabylie du Djurdjura, pic
Hydrogonium bolleanum). According to Ros et al. (2013), its
Belloua; Monts de Ksour, djebel Mzi; Ouarsenis, Téniet el Had. identity has not been revised because the material has not
Jelenc (1955a, 1966, 1967): Kabylie du Djurdjura dans le massif been traced. The localities reported by Jelenc (1955a) were:
de l’Akfadou au Camping Blank, 1500 m. Jelenc (1956, 1967): Cascade du Khémis à Mazer, 1230 m; Monts de Tlemcen;
Atlas Mitidjien, Massif du Tessala, pic Mouzaïa. Jelenc (1967): Monts des Ksour, djebel Mekter au ravin de Aïn ed Diss
Massif de l’Aurès, Ras el Ma; Massif du Tessala, pic de Tafaraoui,
600 m. Ros et al. (1990): Texenna’s wood, 700 m. Ahayoun et al. Foukania, 1400 m.
(2007): Atlas de Blida. Boukhatem et al. (2017): El-Kala National
Park, Tonga lake watershed. Mazari & Boulaacheb (2023): Setif,
Megriss Mountain. Very common. Hydrogonium croceum (Brid.) Jan Kučera

Barbula crocea (Brid.) F.Weber & D.Mohr

*Hydrogonium arcuatum (Griff.) Wijk & Margad. Nomenclature follows Kučera et al. (2013).
Barbula arcuata Griff. Records. — Bescherelle (1882), Jelenc (1955a): Bône. Jelenc
(1955a): Philippeville. Rare.
Nomenclature follows Kučera et al. (2013).

Records. — Boukhatem et al. (2017): El-Kala National Park, Hydrogonium orientale (F.Weber) Jan Kučera
Tonga lake watershed.
Barbula indica (Hook.) Spreng.
Comment Nomenclature follows Kučera et al. (2013).
Doubtfully present in Algeria. This American species was pre-
Records. — Geheeb (1879), Bescherelle (1882): Cascade du Rimel,
viously reported from Egypt by Shabbara (2006). As stated by prope Constantine Algeriae. Bescherelle (1882): Cascades des gorges
Ros et al. (2013), its presence in the Mediterranean Region de la Chiffa; Constantine, bords d’une source thermale. Bescherelle
is unlikely and requires verification. (1882), Jelenc (1955a): Alger. Corbière (1904): Constantine, sources

82 CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5)

 Revisiting the Bryophyte Checklist of Algeria

thermales de Sidi M’Cid. Corbière (1905): Environs de Constan- Hymenoloma crispulum (Hedw.) Ochyra
tine, Moulin Laire. Corbière (1905), Jelenc (1955a): Environs de
Constantine, ravin du Rummel; Environs de Constantine, ruisseau Records. — Bescherelle (1882): Djebel Aurès. Jelenc (1953, 1955a,
près Salah Bey. Jelenc (1949, 1967): Haute plaine de Tlemcen, Aïn 1967): Aurès, flanc nord du Chélia, au Ras Keltoum, 2200 m.
Defla, 620 m; Haute plaine de Tlemcen, oued Saf-Saf, vers 600 m; Jelenc (1953, 1967): Aurès, forêt de Sgag, 1700 m. Boukhatem
Monts de Tlemcen, cirque d’El Ourit. Jelenc (1955a): Gorges de et al. (2017): El-Kala National Park, Tonga lake watershed. Rare.
la Chiffa, ruisseau des Singes. Boukhatem et al. (2017): El-Kala
National Park, Tonga lake watershed. Frequent.
Hymenostomum subrostellatum Schimp. ex Besch.

Hygroamblystegium fluviatile (Hedw.) Loeske Records. — Bescherelle (1882), Jelenc (1955a): Philippeville.

Records. — Jelenc (1955a): Blida; Gorges de la Chiffa; Tlemcen.

Rare. Comment
It was described from Algeria by Bescherelle (1882), but
according to Ros et al. (2013), the type has not been traced
Hygroamblystegium tenax (Hedw.) Jenn. and its identity has not been studied; if it merits recogni-
Records. — Bescherelle (1882): Oued Sidi-Kebir. Bescherelle tion, then a new combination in the genus Weissia Hedw.
(1882), Jelenc (1955a): Alger. Braun-Blanquet & Maire (1924), Je- may be required.
lenc (1955a): Tlemcen. Jelenc (1949, 1967): Vallée de l’Isser, lieu dit
“El Kasbat”, près de Lamoricière, 500 m. Jelenc (1955a): Monts de
Batna, djebel Touggour, 1300 m. Boukhatem et al. (2017): El-Kala
National Park, Tonga lake watershed. Rare. Hymenostylium recurvirostrum (Hedw.) Dixon

Records. — Jelenc (1951, 1955a): Monts de Tlemcen, Lalla Setti,

1000 m. Boukhatem et al. (2017): El-Kala National Park, Tonga
Hygroamblystegium varium (Hedw.) Mönk. lake watershed. Rare.

Records. — Litschauer (1904), Jelenc (1955a): Kabylie du Djurd-

jura, col de Tirourda, 1700 m. Jelenc (1955a): Alger. Rare. Hypnum cupressiforme Hedw. var. cupressiforme

Records. — Bescherelle (1882): Kabylie; La Galite. Bescherel-

*Hylocomiadelphus triquetrus (Hedw.) Ochyra & Stebel le (1882), Jelenc (1955a): Alger, petit Atlas; Édough; La Calle;
Philippeville. Gandoger (1883), Trabut (1888): sine loco. Cor-
Rhytidiadelphus triquetrus (Hedw.) Warnst. bière (1889): Blida. Corbière (1904), Jelenc (1955a): Gorges
Nomenclature follows Ochyra & Stebel (2008) and Ignatov et al. de la Chiffa, ruisseau des Singes et Camp des Chênes. Corbière
(2019). (1904, 1905), Jelenc (1955a): Constantine, bois du djebel Ou-
asch, 1100 m. Litschauer (1904): Chiffaschlucht und im Tal
Records. — Bescherelle (1882), Jelenc (1955a): Kabylie du Djurd- des Qued-Kebir bei Blidah, 300 m. Corbière (1905): Environs
jura, Tizi Ouzou; Tlemcen. Trabut (1888): sine loco. Jelenc (1955a): de Constantinte, Les Pins. Corbière (1905), Jelenc (1955a):
Kabylie du Djurdjura, Aït Idjer. Environs de Constantinte, Sidi M’Cid. Camus (1906), Jelenc
(1955a): Oran. Lapie (1909), Kabylie du Djurdjura. Feldmann
Comment (1947), Jelenc (1955a): Kabylie de Collo, Baie de Tamanart.
Jelenc (1949, 1967): Monts de Tlemcen, pentes du djebel el
Doubtfully present in Algeria. The presence of this Circumpo- Béniane, 1050 m, dans la “Forêt des Pins”; Monts de Tlemcen,
lar Boreo-temperate species (Blockeel et al. 2014b) in Algeria Forêt de Zariffet, berges de l’oued Zarifète, 1100 m; Monts de
is unlikely. Given the challenges in locating the collections, Tlemcen, Koudiat d’Hafir, 1250 m. Jelenc (1953, 1967): Atlas
this species could only be retained in the Algerian catalogue de Blida, ravin Bleu de Chréa. Jelenc (1954b, 1967): Atlas mitid-
upon the discovery of new specimens. jien, föret d’El Aïoun, 500 m; Massif du Bou Zegza, gorges de
Keddara, 400 m. Jelenc (1955a): Alger, Tixérain; Aumale; Aurès;
Djidjelli; Kabylie du Djurdjura, Akfadou; Kabylie du Djurdjura,
Beni Khalfoun; Kabylie du Djurdjura, Fort National; Kabylie du
*Hylocomium splendens (Hedw.) Schimp. Djurdjura, Izer Zouggaren; Kabylie du Djurdjura, Tala Ghizan;
Ouarsenis, Téniet el Had; Oued el Kébir. Jelenc (1956, 1967):
Records. — Bescherelle (1882), Jelenc (1955a): Kabylie du Djurd- Atlas Mitidjien, pic Mouzaïa. Ros et al. (1990): Texenna’s wood,
jura, Tizi Ouzou. 700 m. Boukhatem et al. (2017): El-Kala National Park, Tonga
lake watershed. Common.

Comment
Doubtfully present in Algeria. The report of this Circumpolar Hypnum cupressiforme var. lacunosum Brid.
Wide-boreal species (Blockeel et al. 2014b) in Algeria is based
on a single specimen collected by Gandoger, and its reliabil- Records. — Litschauer (1904), Jelenc (1955a): Atlas de Blida,
oued el Kebir, 200 m; Fort National und Michelet, 1000-1200 m.
ity is doubtful. Considering the difficulties encountered in Doignon (1953): sine loco. Jelenc (1967): Oran, Batterie Espagnole;
locating this collection, the species can only be included in Massif de l’Ouarsenis, Téniet el Had. Mazari & Boulaacheb (2023):
the Algerian catalogue upon the discovery of new specimens. Setif, Megriss Mountain. Rare.

CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5) 83

 Ros R. M. et al.

Hypnum cupressiforme var. subjulaceum Molendo Isothecium myosuroides Brid.

Records. — Litschauer (1904), Jelenc (1955a): Atlas de Blida, Records. — Bescherelle (1882): Kabylie des Babor, Tababor; Oued
oued el Kebir, 200 m. Very rare. Afziz. Feldmann (1947), Jelenc (1955a): Kabylie de Collo, sommet
du djebel Goufi, versant Nord, 1000 m. Jelenc (1955a): Atlas de
Blida; Djebel Chenoua; Tlemcen. Rare.
Hypnum jutlandicum Holmen & E.Warncke

Records. — Corbière (1889), Jelenc (1955a): Blida. Jelenc (1956, Kindbergia praelonga (Hedw.) Ochyra
1967): Massif du Bou Zegza, dans les gorges de Keddara. Jelenc
(1967): Kabylie du Djurdjura, Tizi Ouzou. Boukhatem et al. (2017): Records. — Bescherelle (1882): Birkadem, sous les pins; Djurjura;
El-Kala National Park, Tonga lake watershed. Mazari & Boulaacheb Édough; Kabylie. Bescherelle (1882), Gandoger (1883), Trabut (1888),
(2023): Setif, Megriss Mountain. Rare. Ahayoun et al. (2007): sine loco. Bescherelle (1882), Jelenc (1955a):
Alger; Kabylie du Djurdjura, Tizi Ouzou; Tixeraïn. Bescherelle (1882),
Jelenc (1955a), Boukhatem et al. (2017): La Calle, bords du lac du
Touga (Tonga). Corbière (1889): Blida, intérieur de la fontaine Aine-
**Hypnum uncinulatum Jur. el-Mudoi, 500 m. Corbière (1905): Environs de Constantine, bords
de l’oued Bou Merzoug; Environs de Constantine, ravin du Rummel.
Corbière (1905), Jelenc (1955a): Environs de Constantine, Pépinière
Comment du djebel Ouach. Lapie (1909): Kabylie du Djurjura. Jelenc (1954b,
Excluded from Algeria. According to Ros et al. (2013), this 1967): Massif du Bou Zegza, gorges de Keddara, 400 m. Jelenc (1955a):
Aurès, djebel Chélia; Kabylie du Djurdjura, Aizer; Kabylie du Djurdjura,
species was erroneously listed by Ros et al. (1999) for Algeria Akfadou; Kabylie du Djurdjura, Azerou Tidjer; Kabylie du Djurdjura,
caused by a misinterpretation of the name Hypnum cupres- Fort National, 1000 m; La Bouzaréa; Philippeville. Jelenc (1956, 1967):
siforme var. uncinatum Boulay, which in fact is a synonym of Massif de la Bouzaréa, près des Bains-Romains. Common.
Hypnum cupressiforme var. cupressiforme and not of Hypnum
uncinulatum.
Leptobryum pyriforme (Hedw.) Wilson

Records. — Jelenc (1955a): Tlemcen. Very rare.

Imbribryum alpinum (Huds. ex With.) N.Pedersen

Records. — Desfontaines (1799): sine loco. Brongniart (1882):

Blidah. Litschauer (1904), Jelenc (1955a): Chiffaschlucht und im Leptodictyum riparium (Hedw.) Wanrst.
Tal des Qued-Kebir bei Blidah, 300 m; Fort National, 1000 m.
Corbière (1905), Jelenc (1955a): Environs de Constantine, djebel Records. — Bescherelle (1882), Jelenc (1955a): Alger. Bescherelle
Ouach. Trabut (1914), Jelenc (1955a): Kabylie du Djurdjura, (1882), Jelenc (1955a), Boukhatem et al. (2017): La Calle, bois
Tirourda. Jelenc (1955a): Alger; Atlas de Blida, Aïn Talazid; marécageux, marais du Tonga; Maison-Carrée. Corbière (1904),
Aurès, djebel Chélia; Djebel Smir; Monts de Tlemcen, forêt Jelenc (1955a): Médéa, bois de Tibarrine. Rare.
d’Hafir. Rare.

Leptodon smithii (Hedw.) F.Weber & D.Mohr

Imbribryum mildeanum (Jur.) J.R.Spence
Records. — Bescherelle (1882): Alger, de Birmandreïs à Birkadem;
Records. — Corbière (1905): Environs de Constantine, Bellevue. Djebel Babor; Kabylie. Bescherelle (1882), Corbière (1904), Jelenc
Jelenc (1955a): Djurdjura; Monts de Constantine, djebel Ouach. (1955a), Ahayoun et al. (2007): Gorges de la Chiffa. Bescherelle (1882),
Mazari & Boulaacheb (2023): Setif, Megriss Mountain. Rare. Jelenc (1955a): Constantine, Hammam Meskoutine; Djebel Thaya;
Kabylie du Djurdjura, Akfadou; La Calle; Philippeville. Gandoger
(1883): sine loco. Corbière (1904), Jelenc (1955a): Monts de Tlemcen,
Sidi-Bou-Médine. Litschauer (1904), Jelenc (1955a): Bône. Corbière
Isopterygiopsis pulchella (Hedw.) Z.Iwats. (1905), Lapie (1909), Jelenc (1955a): Kabylie du Djurdjura, Beni Khal-
foun. Jelenc (1949): escarpements du djebel el Béniane. Jelenc (1951,
Records. — Jelenc (1955a): Djurdjura, Ifri n’Ouadnefel. Very rare. 1967): Monts de Tlemcen, El Aguiba, 900 m. Jelenc (1955a): Alger,
La Bouzaréa; Atlas de Blida; Aurès; Corso; Djebel Mouzaïa; Édough;
Kabylie des Babor, cap Aokas; Kabylie du Djurdjura, Toudja; Oued
Djemaa. Jelenc (1955a, 1967): Kabylie de Collo, djebel Goufi; Jelenc
Isothecium alopecuroides (Lam. ex Dubois) Isov. (1956, 1967): Atlas Mitidjien, près du Col des Deux-Bassins; Massif
du Bou Zezga, gorges de Keddara, 400 m. Jelenc (1966, 1967): Monts
de Constantine, djebel Ouach, 1000 m. Common.
Records. — Bescherelle (1882): Djurjura. Bescherelle (1882), Jelenc
(1955a): Kabylie du Djurdjura, Tizi Ouzou. Bescherelle (1882),
Jelenc (1955a, 1967): Djebel Goufi. Lapie (1909): Kabylie du Dju-
rjura. Jelenc (1955a): Atlas de Blida; Boghar; Djidjelli; Kabyblie du Lescuraea incurvata (Hedw.) E.Lawton
Djurdjura, Aït Idjer; Kabyblie du Djurdjura, Akfadou; Kabylie du
Djurdjura, Izer Zouggaren. Jelenc (1955a), Ahayoun et al. (2007): Pseudoleskea perraldieri Besch.
Kabylie du Djurdjura, Tala Ghizan. Boukhatem et al. (2017): El-Kala
National Park, Tonga lake watershed. Rare. Nomenclature follows Ellis et al. (2017).

84 CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5)

 Revisiting the Bryophyte Checklist of Algeria

Records. — Bescherelle (1882), Jelenc (1955a): Kabylie des Babor, Nomenclature follows Lara et al. (2016).
Djebel Tababor. Lapie (1909): Kabylie du Djurjura. Jelenc (1953,
1955a, 1967): Kabylie du Djurdjura, massif de l’Aizer, flanc nord Records. — Bescherelle (1882): Petit Atlas. Brongniart (1882): Bli-
du djebel Tachgagalt, 1950-2000 m. Jelenc (1955a): Kabylie du dah. Corbière (1889), Jelenc (1955a): forêt des Beni-Salah, 1500 m.
Djurdjura, Aït Ali; Monts de Tlemcen, forêt d’Hafir. Rare. Corbière (1904, 1905), Jelenc (1955a, 1966, 1967): Environs de
Constantine, djebel Ouasch, 1000-1100 m. Camus (1906), Jelenc
(1955a): Oran. Jelenc (1953, 1967): Massif de l’Ouarsenis, cédraie
de Téniet el Had, 1500 m. Jelenc (1955a): Kabylie du Djurdjura,
Lescuraea mutabilis (Brid.) Lindb. ex I.Hagen Aït Idjer; Kabylie du Djurdjura, Akfadou; Kabylie du Djurdjura,
Fort National, 1000 m; Monts de Batna, djebel bou Arif; Monts de
Records. — Bescherelle (1882), Jelenc (1955a): Atlas, sur les cèdres; Constantine. Jelenc (1955a), Ahayoun et al. (2007): Atlas de Blida.
Kabylie, Tizi-Ouzou, sommet du Belloua. Gandoger (1883): sine Jelenc (1956, 1967): Massif du Tessala, pic Tessala, 1000 m. Jelenc
loco. Jelenc (1955a): Alger; Atlas de Blida. Rare. (1967): Kabylie du Djurdjura, Aizer au pic des Cèdres, 1880 m;
Massif de l’Ouarsenis, pic Sidi Abdelkader, 1500 m. Boukhatem
et al. (2017): El-Kala National Park, Tonga lake watershed. Mazari &
Leskea polycarpa Hedw. Boulaacheb (2023): Setif, Megriss Mountain. Frequent.
Records. — Jelenc (1955a): Alger. Very rare.
Lewinskya breviseta (F.Lara, Garilleti & Mazimpaka)
F.Lara, Garilleti & Goffinet
Leucodon sciuroides (Hedw.) Schwägr.
Orthotrichum speciosum var. brevisetum F.Lara, Garilleti & Mazimpaka
Records. — Bescherelle (1882): Djebel Édough; Djurjura; La
Calle; Le petit Atlas; Médéah; Philippeville. Bescherelle (1882), Nomenclature follows Lara et al. (2016).
Jelenc (1955a): Aurès; Djebel Babor; Kabylie du Djurdjura, Ak- Records. — Draper et al. (2003): Jijel, djebel Tababor. Mazari &
fadou; Oran, partie méridionale. Brongniart (1882): Blidah. Trabut Boulaacheb (2023): Setif, Megriss Mountain. Rare.
(1887a): D’Oran à Mécheria. Corbière (1889), Jelenc (1955a):
Atlas de Blida, sommet de l’Abd-el-Kader. Litschauer (1904):
Col de Tirourda, 1700 m. Litschauer (1904), Jelenc (1955a): Comment
Fort National nach Michelet, 1000 m. Corbière (1905), Jelenc
(1955a, 1966, 1967): Monts de Constantine au djebel Ouach, Erroneously, Ros et al. (2013) did not report this taxon from
1000 m. Lapie (1909): Kabylie du Djurjura, Beni Khalfoun. Algeria.
Jelenc (1949, 1967): Haute-Plaine de Sétif, Ain Trich, 1100 m;
Monts de Tlemcen, djebel el Beniane, 1150 m. Jelenc (1953,
1967): Atlas de Blida, Cèdraies du ravin Bleu à Chrèa; Kabylie Lewinskya fastigiata (Bruch ex Brid.)
du Djurdjura, massif de l’Aizer, forêt de Ait Ali, 1600 m; Massif
de l’Ouarsenis, flanc nord du Pic Sidi Abdelkader, 1500 m. Jel- Vigalondo, F.Lara, Garilleti & Goffinet
enc (1953, 1967), Ahayoun et al. (2007): Massif de l’Ouarsenis,
cédraie de Téniet el Had, 1500 m. Jelenc (1954b, 1967): Massif Orthotrichum fastigiatum Bruch ex Brid.
de l’Akfadou, camping Blank, 1500 m. Jelenc (1955a): Alger;
Kabylie de Collo, djebel Goufi; Kabylie des Babor, Dra el Arba; Nomenclature follows Vigalondo et al. (2019).
Kabylie du Djurdjura, Aït Ali; Kabylie du Djurdjura, Aït Idjer; Records. — Bescherelle (1882), Jelenc (1955a): Sommet de l’Atlas,
Kabylie du Djurdjura, Aït Ouaban; Kabylie du Djurdjura, Dra el Atlas de Blida. Very rare.
Mizan; Kabylie du Djurdjura, Dra Inguel; Kabylie du Djurdjura,
Tala Ghizan; Massif de la Mahouna, Serdj el Aouda; Ouarsenis,
djebel Doui. Jelenc (1956): Atlas Mitidjien. Jelenc (1967): At- Comment
las Mitidjien, col des Deux-Bassins, 850 m; Kabylie de Collo.
Boukhatem et al. (2017): El-Kala National Park, Tonga lake This species was previously considered a synonym of Lewin-
watershed. Common. skya affinis (synonym: Orthotrichum affine Schrad. ex Brid.),
and so was considered by Ros et al. (1999, 2013) but it was
reinstated by Vigalondo et al. (2019).
Lewinskya acuminata
(H.Philib.) F.Lara, Garilleti & Goffinet
Lewinskya rupestris
Orthotrichum acuminatum H.Philib.
(Schleich. ex Schwägr.) F.Lara, Garilleti & Goffinet
Nomenclature follows Lara et al. (2016).
Orthotrichum rupestre Schleich. ex Schwägr.
Records. — Corbière (1904, 1905), Jelenc (1955a): Environs de
Constantine, bois du djebel Ouasch, 1100 m. Jelenc (1955a): Kaby- Orthotrichum rupestre var. vulgare Venturi
lie. Mazari & Boulaacheb (2023): Setif, Megriss Mountain. Rare.
Nomenclature follows Garilleti et al. (2007) and Lara et al. (2016).
Records. — Bescherelle (1882): Blidah, troncs des chênes verts;
Lewinskya affinis Djebel Édough, sur le chêneliège; Kabylie. Bescherelle (1882),
(Schrad. ex Brid.) F.Lara, Garilleti & Goffinet Jelenc (1955a): Aurès; Djebel Tababor; Massif de la Mahouna, Serdj
el Aouda. Corbière (1889), Jelenc (1953): Pic Sidi Abdelkader,
Orthotrichum affine Schrad. ex Brid. 1500 m. Camus (1906): Oran. Jelenc (1951, 1955a, 1967): Monts

CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5) 85

 Ros R. M. et al.

de Tlemcen, forêt de Tessera Mramet, 1400 m. Jelenc (1953): Records. — Philibert (1880b): Forêt de Ceiba. Bescherelle (1882),
Djurdjura, Aizer, au sommet du pic des Cédres, 1880 m. Jelenc Jelenc (1955a): Djebel Mouzaïa, 1100 m. Jelenc (1955a): Djurdjura;
(1953, 1955a, 1967): Kabylie des Babor, flanc nord du djebel Ba- Gorges de la Chiffa; Kabylie des Babor, Tababor. Rare.
bor, 1700 m. Jelenc (1953, 1967): Massif de l’Ouarsenis, cédraie
de Téniet el Had, 1500 m. Jelenc (1955a): Atlas de Blida; Kabylie
du Djurdjura, Akfadou; Kabylie du Djurdjura, col de Tirourda,
1700 m; Kabylie du Djurdjura, djebel Afroun; Kabylie du Djurd- Microbryum curvicollum (Hedw.) R.H.Zander
jura, Tala Ghizan; Monts de Tlemcen, forêt d’Hafir. Jelenc (1955a),
Ahayoun et al. (2007): Tiaret. Jelenc (1967): Massif du Bou Zezga, Records. — Montagne (1856), Bescherelle (1882), Jelenc (1955a):
gorges de Keddara, 400 m. Bizot (1973): Djebel Belezma à 15 km Blidah, bord des haies. Jelenc (1949, 1955a): Monts de Tlemcen,
à l’ouest de Batna. Mazari & Boulaacheb (2023): Setif, Megriss djebel el Beniane, 900 m. Guerra et al. (1991), Ros et al. (2005):
Mountain. Common. sine loco. Rare.

Lewinskya speciosa (Nees) F.Lara, Garilleti & Goffinet Microbryum davallianum (Sm.) R.H.Zander

Orthotrichum speciosum Nees Records. — Bescherelle (1882): Constantine; Coteaux de Bab-Azoun,

de l’Agha; Mustapha. Bescherelle (1882), Corbière (1904), Jelenc
Nomenclature follows Lara et al. (2016). (1955a): Alger; Oran. Bescherelle (1882), Jelenc (1955a): Blidah; La
Calle. Trabut (1887a): D’Oran à Mécheria. Corbière (1904), Jelenc
Records. — Corbière (1889), Jelenc (1955a): Atlas de Blida, som- (1955a): Orléansville. Corbière (1905), Jelenc (1955a): Environs de
met du pic Abd el Kader. Jelenc (1948): Atlas de Blida; Djurdjura. Constantine, près de l’oued Melah. Braun-Blanquet & Maire (1924),
Jelenc (1955a): Kabylie du Djurdjura, Aït Ouaban. Bizot (1973): Jelenc (1955a): Tlemcen. Jelenc (1949, 1967): Haute plaine de Tlem-
Djebel Belezma à 15 km à l’ouest de Batna. Rare. cen, Berges du chabet bel Abbès, 775 m; Haute plaine de Tlemcen,
Vallée de l’oued Safsaf au pied du djebel el Melah, 650 m; Vallée de
l’Isser, source thermale de Sidi Abdelly, 375 m. Jelenc (1951), Jelenc
Lewinskya striata (Hedw.) F.Lara, Garilleti & Goffinet (1967): Haute plaine de Tlemcen, 600 m. Jelenc (1955a): Bône;
Kabylie du Djurdjura, Fort National; Monts de Constantine, oued
Orthotrichum striatum Hedw. bou Merzoug. Ros et al. (1990): Ouled Mimoum. Boukhatem et al.
(2017): El-Kala National Park, Tonga lake watershed. Frequent.
Orthotrichum biocarpum Bruch & Schimp., invalid, orthographic
variant of O. leiocarpum Bruch & Schimp.
Comment
Nomenclature follows Vitt (1973) and Lara et al. (2016). No varieties have been recognized within M. davallianum, fol-
Records. — Bescherelle (1882): Atlas, sur les chênes-verts. Cor-
lowing the criterium of Ros et al. (1996) as the morphological
bière (1889), Jelenc (1955a): Atlas de Blida, forêt des Beni Salah, characteristics of M. davallianum var. conicum (Schleich. ex
1500 m. Corbière (1905), Jelenc (1955a): Environs de Constantine, Schwägr.) R.H.Zander and M. davallianum var. commutatum
djebel Ouach. Jelenc (1953, 1967): Massif de l’Aurès, forêt de Sgag, (Limpr.) R.H.Zander accepted in Hodgetts et al. (2020)
1700 m. Jelenc (1954b, 1967): Monts de Tlemcen, forêt(fort) de and described in detail by Pilkington (2022) for Britain and
Zariflet, près de l’Aïn Defla, 1200 m. Jelenc (1955a): Kabylie du
Djurdjura, Tala Ghizan. Mazari & Boulaacheb (2023): Setif, Me- Ireland overlap to a great extent in Mediterranean samples.
griss Mountain. Rare.

Microbryum floerkeanum (F.Weber & D.Mohr) Schimp.

*Loeskeobryum brevirostre (Brid.) M.Fleisch. Records. — Bescherelle (1882), Jelenc (1955a): Philippeville.
Very rare.
Records. — Bescherelle (1882), Jelenc (1955a): Kabylie du Djurd-
jura, Taguemmount Azouz. Very rare.
Microbryum rectum (With.) R.H.Zander
Comment Records. — Bescherelle (1882): Agha-Supérieur; Alger; Birkadem,
Doubtfully present in Algeria. The report of this European cimetière israélite. Bescherelle (1882), Jelenc (1955a): Kabylie du
temperate species (Blockeel et al. 2014b) in Algeria is based Djurdjura, pic Belloua, 1450 m. Gandoger (1883): sine loco. Jelenc
on a single specimen collected by Gandoger, and its reliabil- (1955a): Kabylie du Djurdjura, Tizi Ouzou. Jelenc (1955b, 1967):
Maison-Carrée. Rare.
ity is doubtful. Considering the difficulties encountered in
locating this collection, the species could only be retained in
the Algerian catalogue upon the discovery of new specimens.
Microbryum starckeanum (Hedw.) R.H.Zander
Records. — Montagne (1838), Bescherelle (1882), Jelenc (1955a),
Metaneckera menziesii (Drumm.) Steere Ahayoun et al. (2007): Alger. Bescherelle (1882): Oran. Besche-
relle (1882), Corbière (1889), Jelenc (1955a): Blidah. Bescherelle
Neckera menziesii Drumm. (1882), Jelenc (1955a): Alma; Bône; Constantine; Djebel Man-
sourah; La Calle; Sommet du Bouzaréah. Trabut (1887a): D’Oran
Nomenclature follows Enroth et al. (2022). à Mécheria. Corbière (1904): Alger, talus dans le faubourg de

86 CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5)

 Revisiting the Bryophyte Checklist of Algeria

Telemly. Corbière (1905): Environs de Constantine, chemin de du Djurdjura, plateau de l’Aizer, 1900 m. Boukhatem et al. (2017):
la tuilerie Moutin; Environs de Constantine, Les Pins; Environs El-Kala National Park, Tonga lake watershed. Rare.
de Constantine, ravin de Constantine; Environs de Constantine,
Salah Bey; Environs de Constantine, Sidi M’Cid. Coppey (1911),
Trabut (1927): Beni-Ounif, Sahara algérien. Jelenc (1951, 1967):
Sahel d’Oran, Massif du Murdjadjo, forêt de M’sila, près de Msa- Myurella tenerrima (Brid.) Lindb.
bia. Jelenc (1955a): Atlas de Blida, oued el Kebir; Corso; Djebel
Ouach; Littoral oranais; Rouiba; Sud oranais, Beni Ounif, à Beni Records. — Bescherelle (1882): Kabylie. Jelenc (1955a): Djurd-
Zireg. Jelenc (1956, 1967): Monts de Tlemcen, vallèe de l’oued jura. Rare.
Isser près de Lamoricière, 750 m. Frahm (1978): Laghouat. Ros
et al. (1990): Dellys, 0-40 m; Ouled Mimoum. Cano et al. (2000):
Grès à Ben Zireg. Ahayoun et al. (2007): Alger, ponite Pescade. Neckera pumila Hedw.
Boukhatem et al. (2017): El-Kala National Park, Tonga lake wa-
tershed. Common. Records. — Feldmann (1947), Jelenc (1955a, 1959): Kabylie de
Collo, pentes Nord du djebel Goufi, 1000 m. Very rare.

Microbryum starckeanum agg. Neotrichostomum crispulum (Bruch) R.H.Zander

Records. — Warnstorf (1916), Jelenc (1955b, 1967): Algier, Sidi Trichostomum crispulum Bruch
Madaus, Grique de la chife. Braun-Blanquet & Maire (1924), Jelenc
(1955a): Tlemcen. Jelenc (1955a): Alger; Atlas de Blida, oued el Nomenclature follows Zander (2023).
Kebir, 200 m; Oran; Pointe-Pescade.
Records. — Bescherelle (1882): Bab-Azoun; Blidah; Pointe-Pes-
Comment cade; Tiaret. Bescherelle (1882), Jelenc (1955a): Alger; Alger, près
de la Masion-Carrée; Bône; Bouzaréah; Kouba, plants d’olivers;
Ros et al. (1996) assigned to this aggregate thee names related La Calle; Philippeville; Stora. Gandoger (1883): sine loco. Julien
to Microbryum starckeanum and M. davallianum whose iden- (1894): Région de Constantine. Corbière (1904), Jelenc (1955a):
tification is difficult to assess because the spore ornamentation Orléansville, bois de la Pépinière. Lapie (1909): Kabylie du Djur-
present intermediate characters between M. davallianum and jura. Jelenc (1949, 1967): Monts de Tlemcen, flanc nord du djebel
M. starckeanum or are completely smooth; also they treated as el Béniane, 1000 m. Jelenc (1951): Monts de Tlemcen. Jelenc
(1951, 1967): Massif du Tessala, pic de Tafaraoui, 500 m. Jelenc
belonging to this aggregate those whose types were destroyed. (1953, 1967): Atlas de Blida, ravin Bleu de Chréa. Jelenc (1955a):
The Algerian reports included here correspond to the names Atlas de Blida, oued el Kebir; Aurès; Constantine; Djebel Chenoua;
Pottia appertii Warnst., Pottia mutica Vent., and Pottia recur- Gorges de la Chiffa, ruisseau des Singes, 300 m; Gue de Constan-
vifolia Warnst. tine; Kabylie du Djurdjura, Aït Khalfoun; Kabylie du Djurdjura,
Dellys. Jelenc (1956, 1967): Massif du Bou Zegza, dans les gorges
de Keddara, 400 m. Jelenc (1966, 1967): Monts de Constantine
au djebel Ouach; Monts de Tlemcen, forêt de Zaarifett, 1100 m.
Microeurhynchium pumilum (Wilson) Ros et al. (1990): Texenna’s wood, 700 m. Ahayoun et al. (2007):
Ignatov & Vanderp. Mont Chanoua. Boukhatem et al. (2017): El-Kala National Park,
Tonga lake watershed. Common.
Records. — Bescherelle (1882), Jelenc (1955a): Djebel Édough,
fond des ravins humides. Lapie (1909): Kabylie du Djurjura. Jelenc
(1953, 1967): Kabylie du Djurdjura, plateau de l’Aizer, 1900 m.
Jelenc (1955a): Alger; Kabylie; Kéddara. Rare. Nogopterium gracile (Hedw.) Crosby & W.R.Buck

Records. — Bescherelle (1882): Atlas, troncs et rochers. Be-

Mnium hornum Hedw. scherelle (1882), Jelenc (1955a): Bône; Gorges de la Chiffa; Kab-
ylie du Djurdjura, Tizi Ouzou; La Calle; Philippeville. Gandoger
(1883): sine loco. Litschauer (1904), Jelenc (1955a): Kabylie du
Records. — Bescherelle (1882), Jelenc (1955a): Djurdjura. Very rare. Djurdjura, Fort National, Michelet, 1000-1200 m; Qued-Kebir
bei Blidah, 200 m. Corbière (1905), Jelenc (1955a): Environs de
Constantine, djebel Ouach; Environs de Constantine, Sidi M’Cid.
Jelenc (1949, 1956, 1967): Monts de Tlemcen, djebel el Béniane,
Mnium marginatum (Dicks. ex With.) P.Beauv. forêt de Zariflet, 1100 m. Jelenc (1955a): Alger; Djidjelli; Kabylie
de Collo, djebel Goufi; Kabylie du Djurdjura, Aizer; Kabylie du
Records. — Lapie (1909): Kabylie du Djurjura. Jelenc (1953, Djurdjura, Port Gueydon; Ouarsenis, djebel Doui; Tlemcen. Ros
1955a, 1967): Kabylie du Djurdjura, massif de l’Aizer flanc nord et al. (1990): Texenna’s wood, 700 m. Ahayoun et al. (2007): Atlas
du djebel Tachgagalt, 2000 m. Rare. de Blida. Boukhatem et al. (2017): El-Kala National Park, Tonga
lake watershed. Common.

Mnium stellare Hedw.

Orthothecium intricatum (Hartm.) Schimp.
Records. — Battandier et al. (1914), Jelenc (1955a): Kabylie du
Djurdjura, Forêt des Aït-Ouaban, groute l’Ifri-bou-Anou. Lapie Records. — Jelenc (1955a): sine loco. Jelenc (1955b, 1967): Ka-
(1909): Kabylie du Djurjura. Trabut (1914), Jelenc (1955a): Kabylie bylie du Djurdjura, Ouadenfej. Boukhatem et al. (2017): El-Kala
du Djurdjura, forêt d’Akfadou. Jelenc (1953, 1955a, 1967): Kabylie National Park, Tonga lake watershed. Rare.

CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5) 87

 Ros R. M. et al.

Orthotrichum anomalum Hedw. Orthotrichum letourneuxii Besch.

Records. — Bescherelle (1882): Atlas; Constantine, rochers du Comment

Rummel et du Chatabah. Bescherelle (1882), Jelenc (1955a): Dje- This species was described from Algeria by Bescherelle (1882)
bel Thaya, rochers; Milah; Tlemcen. Corbière (1904): Tlemcen,
rochers sur les hauteurs, entre Sidi-Bou-Médine et les cascades.
from Djebel Thaya and later compiled in Jelenc (1955a).
Corbière (1905), Jelenc (1955a): Environs de Constantine, Les According to Lewinsky (1993), the type cannot be located,
Pins; Environs de Constantine, route de la Corniche; Environs de and it is impossible to conclude from the protologue whether
Constantine, Sidi M’Cid. Thériot & Trabut (1930), Jelenc (1955a), it is an Orthotrichum or not.
Ros & Cano (2008): Babor. Jelenc (1949): Monts de Tlemcen, forêt
de Tessera Mramet, 1400 m. Jelenc (1955a): Atlas de Blida, oued
el Kébir, 200-300 m; Constantine; Djebel Chenoua; Gorges de la
Chiffa; Kabylie du Djurdjura, Aït Idjer; Kabylie du Djurdjura, Ak- Orthotrichum macrocephalum
fadou; Kabylie du Djurdjura, Fort National; Kabylie du Djurdjura, F.Lara, Garilleti & Mazimpaka
Tirourda; Lambèse. Jelenc (1956): Massif du Bou Zegza, dans les
gorges de Keddara. Bizot (1973): Djebel Belezma à 15 km à l‘ouest Records. — Mazari et al. (2024): Sétif, Djebel Megriss Mountain,
de Batna. Boukhatem et al. (2017): El-Kala National Park, Tonga Ain Guelou, 36°21’40.0”N, 5°20’19.8”E, c. 1108 m; Sétif, Djebel
lake watershed. Common. Megriss Mountain, Oued El Bordj, 36°20’5.8”N, 5°20’31.5”E,
c. 1554 m. Rare.

Orthotrichum cupulatum Hoffm. ex Brid.

Orthotrichum patens Bruch ex Brid.
Records. — Bescherelle (1882), Braun-Blanquet & Maire (1924),
Records. — Jelenc (1955a): Kabylie du Djurdjura, Tala Ghizan.
Jelenc (1955a): Tlemcen. Bescherelle (1882), Jelenc (1955a): Al-
Very rare.
ger; Atlas de Blida, Beni Salah; Constantine, ravin du Rummel;
Djebel Thaya; Kabylie du Djurdjura, col de Tirourda; Médéah.
Julien (1894): Région de Constantine. Corbière (1904), Jelenc
(1955a): Sétif, 1100 m. Corbière (1905): Environs de Constan- Orthotrichum philibertii Venturi
tine, Les Pins; Environs de Constantine, route de la Corniche;
Environs de Constantine, Sidi M’Cid. Jelenc (1953, 1967): Massif Records. — Blockeel et al. (2003): Forêt des Senhadja, near Dem
du Bélézma, cédraies de l’oued Chaba, 1400 m. Jelenc (1955a): El Begrat, south of Skikda. Very rare.
Aumale; Djebel Chenoua; Djebel Doui; Kabylie du Djurdjura,
djebel Afroun; Lambèse; Ouarsenis. Jelenc (1956, 1967): Atlas
Mitidjien, pic Mouzaïa; Massif du Dahra, à Lapasset. Jelenc Orthotrichum pumilum Sw. ex anon.
(1967): Monts de Tlemcen, forêt de Tessera Mramet, 1400 m.
Boukhatem et al. (2017): El-Kala National Park, Tonga lake wa- Records. — Corbière (1905): Constantine. Corbière (1905),
tershed. Common. Jelenc (1955a): Environs de Constantine, Pépinière du djebel
Ouach; Environs de Constantine, Sidi M’Cid; Environs de Con-
stantine, sur des arbres au Hamma; Environs de Constantine,
sur des aulnes au bord du Rummel. Jelenc (1955a): Kabylie du
Orthotrichum cupulatum var. riparium Huebener Djurdjura, Fort National. Mazari & Boulaacheb (2023): Setif,
Megriss Mountain. Rare.
Records. — Bescherelle (1882), Jelenc (1955a): Constantine.
Bescherelle (1882), Jelenc (1955a, 1955b): Batna. Jelenc (1955a):
Aurès. Jelenc (1955b, 1967): Monts de Constantine. Mazari &
Boulaacheb (2023): Setif, Megriss Mountain. Rare. Orthotrichum scanicum Grönvall

R ecords . — Mazari & Boulaacheb (2023): Setif, Megriss

Mountain, Ain Guelou, 36°21’40.0”N, 5°20’19.8”E, c. 1108 m.
Orthotrichum diaphanum Brid. Very rare.

Records. — Bescherelle (1882): Constantine. Bescherelle (1882),

Jelenc (1955a): Alger, sur les cactées; Bouzaréah; La Calle; Mas- Orthotrichum schimperi Hammar
cara; Philippeville. Trabut (1887a): D’Oran à Mécheria. Corbière
(1889): Blida. Corbière (1904), Jelenc (1955a): Médéa; Sétif, Records. — Bescherelle (1882), Jelenc (1955a): sine loco. Cor-
1060 m. Corbière (1905): Environs de Constantine, Bellevue; bière (1904): Médéa. Corbière (1904), Jelenc (1955a): La Calle;
Environs de Constantine, Chettabah; Environs de Constantine, Tlemcen, sur tronc d’olivier à Sidi-Bou-Médine. Camus (1906),
Menidj; Environs de Constantine, Sidi M’Cid. Corbière (1905), Jelenc (1955a): Oran. Jelenc (1955a): Kabylie du Djurdjura, Fort
Jelenc (1955a): Environs de Constantine, djebel Ouach. Jelenc National. Rare.
(1949, 1967): Massif du Murdjadjo (Oran), Ravin de l’oued Tam-
mermouth; Monts et Haute-Plaine de Tlemcen. Jelenc (1955a):
Atlas de Blida; Aumale; Djebel Smir; Kabylie du Djurdjura, col
des Aït Aïcha; Kabylie du Djurdjura, Fort National; Tixeraïn; Orthotrichum stramineum Hornsch. ex Brid.
Tlemcen, 1000 m. Jelenc (1956, 1967): Sahel d’Alger, environs
d’El Biar. Mazari & Boulaacheb (2023): Setif, Megriss Moun- Records. — Trabut (1914), Jelenc (1955a): Kabylie du Djurdjura,
tain. Common. forêt d’Akfadou. Very rare.

88 CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5)

 Revisiting the Bryophyte Checklist of Algeria

Orthotrichum tenellum Bruch ex Brid. orientale, Adghar Amellal. Brongniart (1882): Blidah. Corbière
(1904), Litschauer (1904): Gorges de la Chiffa, ruisseau des Singes.
Records. — Bescherelle (1882), Jelenc (1955a): Alger; Kabylie du Corbière (1904), Jelenc (1955a): Tlemcen. Litschauer (1904), Jelenc
Djurdjura, Tizi Ouzou; La Calle; Oued-el-Aneb, près Sidi-el-Akhdar; (1955a): Tal des Qued-Kebir bei Blidah, 300 m. Lapie (1909): Kabylie
Philippeville. Gandoger (1883): sine loco. Corbière (1904), Jelenc (1955a): du Djurjura. Jelenc (1951, 1967): Monts de Tlemcen, cirque d‘El
Médéa. Corbière (1905): Constantine; Environs de Constantine, Le Ourit. Jelenc (1955a): Dra Inguel; Kabylie des Babor, Aït Embarek;
Hamma; Environs de Constantine, Sidi M’Cid. Corbière (1905), Jelenc Kabylie du Djurdjura, Aït Ali; Kabylie du Djurdjura, Aït Daoud;
(1955a): Environs de Constantine, djebel Ouach. Jelenc (1951, 1967): Kabylie du Djurdjura, Tala bou Idhen; Kabylie du Djurdjura, Tala
Monts de Tlemcen, forêt de Tessera Mramet, 1400 m. Jelenc (1955a): Ghizan. Jelenc (1967): Kabylie des Babor, djebel Babor, 1300 m;
Édough; Khodjaberry; La Réghaia; Tlemcen, 1000 m. Frequent. Massif de l‘Aurés, Ain Mimoun; Massif de l‘Aurès, djebel Chélia,
1500 m; Massif de l‘Aurès, djebel Pharaoun. Frequent.

Orthotrichum urnigerum Myrin

Palustriella falcata (Brid.) Hedenäs
Records. — Jelenc (1955a): Atlas de Blida; Duperré; Kabylie du
Djurdjura, Tala Ghizan; Ouarsenis, djebel Doui. Rare. Records. — Bescherelle (1882): Atlas, commun dans les ruisse-
aux; Djurjura. Corbière (1889), Jelenc (1955a): Blida, gorges de la
Chiffa. Litschauer (1904), Jelenc (1955a): Kabylie du Djurdjura, col
Oxyrrhynchium hians (Hedw.) Loeske de Tirourda, 1700 m. Jelenc (1953): Aurès, flanc nord du Chélia,
1500 m; Babor, flanc nord du Babor, 1300 m. Jelenc (1954b, 1967):
Records. — Jelenc (1949, 1955a, 1967): Monts de Tlemcen, Mazer. Kabylie du Djurdjura, lac Goulmine, 1600 m. Jelenc (1955a): Atlas
Jelenc (1953, 1967): Kabylie du Djurdjura, massif de l’Aizer, 1900 m. de Blida, Aïn Talazid; Kabylie du Djurdjura, Aït Daoud; Kabylie du
Jelenc (1954b, 1967): Monts de Saida, cascades de Tifrit; Sahel d’Alger, Djurdjura, Tala Ghizan. Boukhatem et al. (2017): El-Kala National
colline du Hamma. Jelenc (1955a): Alger, La Bouzaréa; Atlas de Blida; Park, Tonga lake watershed. Frequent.
Constantine, oued bou Merzoug; Édough; Kabylie du Djurdjura,
Tizi Ouzou. Jelenc (1956): Fontaine du Musée des Beaux-Art. Jelenc
(1956, 1967): Sahel d’Alger, dans le ravin du Frais-Valllon. Boukhatem
et al. (2017): El-Kala National Park, Tonga lake watershed. Frequent. Philonotis caespitosa Jur.

Records. — Pierrot (2000): Environs de Blida, gorges de l’oued

Comment el Kébir. Boukhatem et al. (2017): El-Kala National Park, Tonga
lake watershed. Rare.
Oxyrrhynchium hians var. vulgare (Boulay) Podp. was reported
in Algeria by Corbière (1904) (sub Eurhynchium praelongum
var. vulgare (Boulay) Limpr.) from Bône, djebel Édough, but Philonotis calcarea (Bruch & Schimp.) Schimp.
Ros et al. (2013) mentioned its presence only in Tunisia. Its
identity remains unchecked at present. Records. — Bescherelle (1882): Atlas; Constantine; Djurjura.
Bescherelle (1882), Litschauer (1904), Jelenc (1955a): Cascades de la
Chiffa. Bescherelle (1882), Jelenc (1955a): Blidah, les Deux-Frères;
Bône; Kabylie des Babor, djebel Tababor; Oran. Trabut (1887a):
Oxyrrhynchium speciosum (Brid.) Warnst. D’Oran à Mécheria. Corbière (1889): Blida, Fontaine-Fraîche.
Corbière (1889), Litschauer (1904), Jelenc (1955a): Blida, l’oued
Records. — Corbière (1905): Environs de Constantine, bords de Sidi-el-Kebir, 200 m. Julien (1894): Région de Constantine. Jelenc
l’oued Bou Merzoug. Feldmann (1947), Jelenc (1955a): Kabylie de (1949, 1967): Monts de Tlemcen, cirque d’el Ourit au pied de la
Collo, vallée de l’oued Elli Ankou, 500-600 m. Jelenc (1949, 1967): cascade supérieure. Jelenc (1951, 1967): Haute plaine de Tlemcen,
Monts de Tlemcen. Jelenc (1955a): Djurdjura, Taourirt Ighil. Boukha- cascades de l’oued bou Ennag à la traversée des collines des Beni
tem et al. (2017): El-Kala National Park, Tonga lake watershed. Rare. Mester, 500 m. Jelenc (1953): Aizer, dans uns source en contre-bas
du chemin Lapie, 1800 m; Aurès, flanc sud du Chélia, dans l’oued
Tacherchar, 1700 m; Bèlezma, suintement d’une source ad Chel-
Oxystegus tenuirostris (Hook. & Taylor) A.J.E.Sm. lala, 1800 m; Flanc nord du djebel Pharaoun, dans les torrents
près d’Aïn Mimoun. Jelenc (1953, 1967): Djurdjura, Dra Inguel,
Records. — Bescherelle (1882), Jelenc (1955a): Kabylie des Babor, au-dessous de Tizi Toumdelt, 1600 m. Jelenc (1954b, 1967): Mas-
djebel Tababor. Very rare. sif du Djurdjura, en contre-bas du lac Goulmine, 1600 m. Jelenc
(1955a): Djebel Dira; Kabylie des Babor, Aït Embarek; Kabylie des
Babor, Sebt des Beni Sliman; Kabylie du Djudjura, Aït Ali; Kabylie
Comment du Djudjura, Aït Daoud; Kabylie du Djudjura, djebel Gouraya;
Kabylie du Djudjura, gorges de l’oued Isser; Kabylie du Djudjura,
The criterion established by Zander & Eckel (2019) was Tirourda; Monts de Constantine. Mazari & Boulaacheb (2023):
followed to classify this species within the genus Oxystegus Setif, Megriss Mountain. Common.
(Limpr.) Hilp., rather than Chionoloma Dixon who advocate
for the maintenance of paraphyletic taxa.
Philonotis capillaris Lindb.
Palustriella commutata (Hedw.) Ochyra Records. — Litschauer (1904), Jelenc (1955a): Fort National und
Michelet, 1000-1200 m. Jelenc (1951, 1967): Monts de Tlemcen,
Records. — Bescherelle (1882): Alger, près des sources; Djurjura; Oued Khémis près du village du Khémis, 800 m. Boukhatem et al.
Sources d’Aïn-Talazid. Bescherelle (1882), Jelenc (1955a): Kabylie (2017): El-Kala National Park, Tonga lake watershed. Rare.

CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5) 89

 Ros R. M. et al.

Philonotis fontana (Hedw.) Brid. Comment

Doubtfully present in Algeria. The report in Algeria of this spe-
Records. — Desfontaines (1799): sine loco. Bescherelle (1882): cies is based on a single specimen collected by A. Letourneux
Rive gauche de l‘Adda. Bescherelle (1882), Jelenc (1955a): Blidah,
sources d‘Aïn-Talasid; Tiaret. Litschauer (1904), Jelenc (1955a):
without indication of locality. Furthermore, its reliability is
Kabylie du Djurdjura, Fort National, Michelet, 1000-1200 m. doubtful as it is a rare species in the Mediterranean Region
Lapie (1909): Kabylie du Djurjura. Jelenc (1951, 1967): Monts (Ros et al. 2013; Sabovljevic 2019). Due to difficulties in
de Tlemcen, cirque d‘El Ourit. Jelenc (1954b, 1967): Grand Atlas finding this collection, the species can only be kept in the
central: refuge d‘Isougane n‘Ouagoun, 3000 m. Jelenc (1955a): Algerian catalogue if new specimens are found.
Acherchour n‘Tensaout; Akfadou; Bérard; Gorges de la Chiffa; Tala
bou Idhen; Tirourda. Boukhatem et al. (2017): El-Kala National
Park, Tonga lake watershed. Frequent.
Plagiomnium affine (Funck) T.J.Kop.)

Philonotis marchica (Hedw.) Brid. Records. — Bescherelle (1882): sine loco. Corbière (1889): Blida.
Dubuis & Faurel (1945), Jelenc (1953, 1955a, 1967): Kabylie du
Djurdjura, plateau de l’Aizer, 1900-2000 m. Jelenc (1953, 1967):
Records. — Bescherelle (1882): Constantine, rochers humides, Atlas de Blida, Cèdraies du ravin Bleu à Chrèa; Djurdjura, Aizer,
aux moulins; Édough, talus de la route Randon. Bescherelle (1882), flanc nord du djebel Tachgagalt, 1950-2000 m. Jelenc (1955a): Atlas
Jelenc (1955a): Atlas de Blida, gorges de la Chiffa. Corbière (1904), de Blida; Aurès, djebel Chélia. Rare.
Jelenc (1955a): Bône, ravin humide aux flancs de d’Édough, 200 m.
Jelenc (1955a): Monts de Constantine. Jelenc (1967): Aizer, 1800 m;
Kabylie du Djurdjura, Yakouren, 1000 m; Lac Goulmine, 1600 m;
Massif de Bélézma, Chellala, 1800 m; Massif de l’Aurès, djebel Plagiomnium cuspidatum (Hedw.) T.J.Kop.)
Chélia, 1700 m; Massif de l’Aurès, djebel Pharaoun, Ain Mimoun;
Massif de l’Aurès, oued Tacherchar. Boukhatem et al. (2017): El-Kala Records. — Jelenc (1955a): Djurdjura. Very rare.
National Park, Tonga lake watershed. Frequent.

Philonotis rigida Brid. Plagiomnium elatum (Bruch & Schimp.) T.J.Kop.

Records. — Bescherelle (1882), Jelenc (1955a): Bône. Litschau- Records. — Jelenc (1955a): Atlas de Blida; Aurès; Djurdjura. Rare.
er (1904), Jelenc (1955a): Kabylie du Djurdjura, Fort National,
1000 m. Feldmann (1947), Jelenc (1955a): Kabylie de Collo, Baie
de Tamanart. Jelenc (1948): Hauts-plateaux, Skourah. Rare.
Plagiomnium rostratum (Schrad.) T.J.Kop.)
Records. — Bescherelle (1882): Atlas. Lapie (1909): Kabylie du
Philonotis tomentella Molendo Djurjura. Jelenc (1955a): Atlas de Blida; Kabylie du Djurdjura,
Dra Inguel. Jelenc (1954b, 1967): Massif du Bou Zezga, gorges de
Records. — Corbière (1905), Jelenc (1955a): Environs de Con- Keddara, 400 m. Boukhatem et al. (2017): El-Kala National Park,
stantine, djebel Ouach. Boukhatem et al. (2017): El-Kala National Tonga lake watershed. Rare.
Park, Tonga lake watershed. Mazari & Boulaacheb (2023): Setif,
Megriss Mountain. Rare.
Plagiomnium undulatum (Hedw.) T.J.Kop.)
**Physcomitrium eurystomum subsp. acuminatum (Bruch & Records. — Bescherelle (1882), Lapie (1909): Djurjura. Bescherelle
Schimp.) Giacom. (1882), Jelenc (1955a): Alger; Bône; Édough; L‘Arba, pied de l‘Atlas.
Brongniart (1882): Blidah. Trabut (1888): sine loco. Jelenc (1949,
Comment 1956, 1967): Monts de Tlemcen, forêt de Zariflet, berges de l‘oued
Excluded from Algeria. This taxon was reported by Bescherelle Zarifète, 1000 m. Jelenc (1949, 1967): Monts de Tlemcen, Koudiat
d‘Hafir, 1250 m. Jelenc (1953, 1967): Atlas de Blida, parois d‘une
(1882) from Const. Sahara Mehounech, 390 m, under the source au-dessous de Chrèa, 1000 m; Kabylie du Djurdjura, Aizer,
name “Physcomitrium acuminatum Schl.”. Jelenc (1955a) while au pied des grandes falaises du chemin Lapie, 1700 m; Massif de
compiling previous northern African reports, considered that l‘Aurès, flanc nord du djebel Chélia, 1900 m. Jelenc (1954b, 1967):
Bescherelle referred to a new undescribed taxon “P. acumina- Monts de Tlemcen, massif de Ghar Rouban, l‘oued Zaouia. Jelenc
tum Besch. non Schleich.” and put it as a synonym of Phy- (1955a): Aïn Taya; Atlas de Blida; Djebel Mouzaïa; Gorges de la
Chiffa; Kabylie du Djurdjura, Acherchour n‘Tensaout; Kabylie du
scomitrium longicollum Trab. (synonymized by Ros & Cano Djurdjura, Yakouren. Jelenc (1955a, 1956): Kabylie du Djurdju-
(2008) with Entosthodon hungaricus). Therefore, it was not ra, massif de l‘Akfadou. Jelenc (1967): Akfadou, fontaine des Ifs,
included in the Algerian checklists of Ros et al. (1999, 2013). 1200 m. Frequent.

*Physcomitrium pyriforme (Hedw.) Hampe Comment

Mnium undulatum var. kabylicum Trab. ex Jelenc nom. inval.
Records. — Desfontaines (1799), Bescherelle (1882), Jelenc descr. gall. was reported by Jelenc (1955a) from Djurdjura.
(1955a): sine loco. According to Ros et al. (2013), its identity has not been revised

90 CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5)

 Revisiting the Bryophyte Checklist of Algeria

since. If it merits recognition, then a new combination under although it was not included in any of the publications of
Plagiomnium undulatum (Hedw.) T.J.Kop. may be required. this author.

Plagiothecium nemorale (Mitt.) A.Jaeger Pogonatum aloides (Hedw.) P.Beauv.

Records. — Bescherelle (1882): Atlas; Kabylie. Bescherelle (1882), Records. — Bescherelle (1882), Jelenc (1955a): Alger; Bône;
Jelenc (1955a): Édough. Trabut (1888): sine loco. Jelenc (1955a): Djurjura; Édough. Litschauer (1904), Jelenc (1955a): Fort
Atlas de Blida; Kabyblie du Djurdjura, Akfadou. Rare. National nach Michelet, 1000-1200 m. Jelenc (1951, 1967):
Monts de Tlemcen, forêt de Zariffet au Koudiat d’Hafir, 1250 m.
Jelenc (1955a): Atlas de Blida; Kabylie du Djurdjura, Akfadou;
Plasteurhynchium meridionale (Schimp.) M.Fleisch. Kabylie du Djurdjura, Tifrit; Kabylie du Djurdjura, Tirourda.
Jelenc (1956, 1967): Kabylie du Djurdjura, massif de l’Akfadou,
Records. — Bescherelle (1882), Jelenc (1955a): Djebel Édough, fontaine des Ifs, 1200 m; Maison forestière de l’Agoulmine Ab-
fond des ravins humides; Kabylie. Rare. erkane. Frequent.

Plasteurhynchium striatulum (Spruce) M.Fleisch. Pogonatum nanum (Hedw.) P.Beauv.

Records. — Jelenc (1955a): Atlas de Blida, oued el Kebir, 200 m. Records. — Bescherelle (1882): Atlas; Djebel Édough. Bescherelle
Very rare. (1882), Lapie (1909): Djurjura. Bescherelle (1882), Jelenc (1955a):
Kabylie du Djurdjura, Tizi Ouzou. Brongniart (1882), Corbière
(1889): Blidah, près la gracière Laval. Litschauer (1904), Jelenc
*Platygyrium repens (Brid.) Schimp. (1955a): Fort National und Michelet, 1000-1200 m. Jelenc (1949,
1967): Monts de Tlemcen, forêt de Zariflet, 1150 m; Jelenc (1955a):
Records. — Bescherelle (1882), Jelenc (1955a): Kabylie, Tizi-Ouzou. Atlas de Blida; Kabylie du Djurdjura, Aït Yahia; La Calle. Jelenc
(1967): Kabylie du Djurdjura, Yakouren, 1000 m; Monts de Tlem-
cen, Hafir. Frequent.
Comment
Doubtfully present in Algeria. The report of this Circumpolar
Temperate species (Blockeel et al. 2014b) in Algeria is based Pohlia annotina (Hedw.) Lindb.
on a single specimen collected by Gandoger, and its reliability
is doubtful. Due to the difficulties in finding this collection, Records. — Bescherelle (1882), Jelenc (1955a): Kabylie, pic du
the species can only be kept in the Algerian catalogue if new Lalla-Khadidja. Boukhatem et al. (2017): El-Kala National Park,
Tonga lake watershed. Rare.
specimens are found.

Pohlia atropurpurea (Wahlenb.) Lindb.

Pleuridium acuminatum Lindb.
Records. — Gandoger (1883): sine loco. Litschauer (1904): Chif-
Records. — Pierrot (2000): Forêt de Baïnem. Boukhatem et al. faschlucht und im Tal des Qued-Kebir bei Blidah, 300 m; Michelet
(2017): El-Kala National Park, Tonga lake watershed. Rare. und Fort National, 1000 m; Vallon bei Alger, 300 m. Braun-Blan-
quet & Maire (1924): Tlemcen. Rare.

Pleuridium subulatum (Hedw.) Rabenh. Comment

The reports included here are those published as Bryum
Records. — Bescherelle (1882): Djebel Beni-Salah. Bescherelle
(1882), Corbière (1889), Jelenc (1955a): Blidah. Bescherelle (1882), atropurpureum in which Wahlenberg (Wahlenb.) appears as
Corbière (1904), Jelenc (1955a): Bouzaréa. Bescherelle (1882), Jelenc the authors. Whereas those in which Bruch & Schimp. are
(1955a): Alger; Birkadem; Bône;La Calle; Philippeville; Tiaret. indicated, are considered as reports of Bryum dichotomum
Corbière (1904), Jelenc (1955a): Kabylie, Fort National, 1000 m, (Bryum atropurpureum Bruch & Schimp. nom. illeg.). Never-
1200 m. Corbière (1905), Jelenc (1955a): Environs de Constantine, theless, as Pohlia atropurpurea is an unfrequent species in the
djebel Ouach, 1000 m. Jelenc (1956, 1967): Plaine de la Mitidja,
forêt de la Règhaïa. Frequent. Mediterranean region, the presence of the species in Algeria
should be confirmed.

*Pleurozium schreberi (Willd. ex Brid.) Mitt.

Pohlia cruda (Hedw.) Lindb.
Records. — Jelenc (1967): sine loco.
Records. — Bescherelle (1882), Jelenc (1955a): Kabylie des Babor,
Comment djebel Tababor. Jelenc (1955a): Aurès. Rare.
Doubtfully present in Algeria. According to Jelenc (1967),
the only report of the species in Algeria was taken from Pohlia elongata Hedw.
the distribution data of the species in Allorge (1955), who
stated that it was based on a record of Feldmann in 1940, Records. — Bescherelle (1882), Jelenc (1955a): sine loco. Very rare.

CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5) 91

 Ros R. M. et al.

Comment Polytrichum strictum Menzies ex Brid.

The only report of this species in Algeria is based on a single
specimen collected by Cordier without indication of locality Records. — Jelenc (1956, 1967): Kabylie du Djurdjura, massif
de l’Akfadou, fontaine des Ifs, 1200 m. Boukhatem et al. (2017):
(Bescherelle 1882). El-Kala National Park, Tonga lake watershed. Rare.

Pohlia melanodon (Brid.) A.J.Shaw Pottiopsis caespitosa (Bruch ex Brid.)

Blockeel & A.J.E.Sm.
Records. — Bescherelle (1882): Alger-Mustapha, rochers humides;
Atlas; Constantine, talus humides du Mansourah; Ravins de Bab- Records. — Bescherelle (1882), Jelenc (1955a): sine loco. Jelenc
el Oued. Bescherelle (1882), Jelenc (1955a): Gorges de la Chiffa, (1966, 1967): Mitidja, forêt de la Réghaia; Monts de Tlemcen à
300 m; Réghaïa. Julien (1894): Région de Constantine. Corbière Lamoricière, 750 m. Ros et al. (1990): Between Mostaganem and
(1904), Jelenc (1955a): Sidi-Bel-Abbès. Corbière (1905): Environs Cap Ivi, Abdelmaimalek-Ramdan beach, 0 m; Chiffa’s Gorge; Djebel
de Constantine, La Pépinière; Environs de Constantine, Le Hamma. Chenoua, 5-40 m; Oued Rhiou. Rare.
Maire (1916), Jelenc (1955a): Sud oranais, le djebel Mzi, 2130 m.
Braun-Blanquet & Maire (1924), Jelenc (1955a): Tlemcen. Jelenc
(1955a): Alger; Atlas de Blida; Constantine; Kabylie du Djurdjura,
djebel Gouraya; Orléansville. Frequent. Pseudephemerum nitidum (Hedw.) Loeske
Records. — Bescherelle (1882), Jelenc (1955a): Bône. Jelenc
(1955a): Alger; Blida; La Calle; Tiaret. Rare.
Pohlia wahlenbergii
(F.Weber & D.Mohr) A.L.Andrews
Pseudocrossidium hornschuchianum
Records. — Bescherelle (1882), Jelenc (1955a): Blidah, cascades de (Schultz) R.H.Zander
l’Atlas. Corbière (1904), Jelenc (1955a): Vallon d’El Biar près Alger,
140 m. Corbière (1905): Environs de Constantine, fossé près les Arcades Records. — Bescherelle (1882): Alger, au-dessus du Fort l‘Emp-
romaines; Environs de Constantine, gue Martin; Environs de Constantine, ereur; Constantine, cimetiére maure. Bescherelle (1882), Jelenc
ruisseau près Salah Bey. Jelenc (1955a): Cherchell; Constantine. Rare. (1955a): Philippeville. Corbière (1904), Jelenc (1955a): Sétif, côteau
au N-E de la ville, 1080 m. Corbière (1905), Jelenc (1955a): Envi-
rons de Constantine, près de l’oued Melah. Jelenc (1955a): Alger;
Polytrichastrum alpinum (Hedw.) G.L.Sm. Maison-Carrée; Rovigo. Ros et al. (1990): Between Mostaganem
and Cap Ivi, Abdelmaimalek-Ramdan beach, 0 m; Oued Rhiou;
Ouled Mimoum. Boukhatem et al. (2017): El-Kala National Park,
Records. — Jelenc (1955a): Kabylie du Djurdjura, Aït Yahia. Very rare.
Tonga lake watershed. Mazari & Boulaacheb (2023): Setif, Megriss
Mountain. Frequent.

Polytrichum commune Hedw.

Pseudocrossidium revolutum (Brid.) R.H.Zander
Records. — Pierrot (2000): Kabylie de Collo, djebel Goufi. Bouk-
hatem et al. (2017): El-Kala National Park, Tonga lake watershed. Rare. Records. — Bescherelle (1882), Jelenc (1955a): Alger; Bouza-
réah; Constantine. Julien (1894): Région de Constantine. Cor-
bière (1905): Environs de Constantine, Les Pins; Environs de
Polytrichum formosum Hedw. Constantine, Mansourah. Jelenc (1955a): Atlas de Blida, oued el
Kebir; Médéa. Bizot (1973): Fontaine Chaude à 30 km NE de
Records. — Bescherelle (1882), Jelenc (1955a): Kabylie du Djurd- Batna. Boukhatem et al. (2017): El-Kala National Park, Tonga
jura, Tala Ghizan. Jelenc (1955a): Kabylie du Djurdjura, Tifrit. Rare. lake watershed. Rare.

Polytrichum juniperinum Hedw.

Pseudoleskeella catenulata (Brid. ex Schrad.) Kindb.
Records. — Bescherelle (1882), Jelenc (1955a): Blidah, petit Atlas;
Djebel Édough, route de Randon. Litschauer (1904), Jelenc (1955a): Records. — Jelenc (1953, 1967): Massif de l’Aurès, forêt de Sgag,
Fort National und Michelet, 1000-1200 m. Corbière (1905), Jelenc 1700 m. Very rare.
(1955a): Environs de Constantine, djebel Ouach. Lapie (1909): Ka-
bylie du Djurjura. Jelenc (1951): Monts de Tlemcen. Jelenc (1955a):
Bône; Kabylie du Djurdjura, Akfadou; Kabylie du Djurdjura, Dje- Pseudoleskeella rupestris
maâ Saharidij; Kabylie du Djurdjura, Port Gueydon. Jelenc (1955a, (Berggr.) Hedenäs & L.Söderstr.
1967): Monts de Tlemcen, Tlemcen, forêt d’Hafir. Boukhatem et al.
(2017): El-Kala National Park, Tonga lake watershed. Frequent. Records. — Jelenc (1955a): sine loco. Very rare.

Polytrichum piliferum Hedw. Comment

The only report of the species in Algeria is based in a single
Records. — Feldmann (1947), Jelenc (1955a): Kabylie de Collo, specimen collected by Allorge without indication of locality
Taoulel, 600 m. Very rare. (Jelenc 1955a).

92 CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5)

 Revisiting the Bryophyte Checklist of Algeria

Pseudorhynchostegiella duriaei from environs de Constantine, Chettabah. Its identity has

(Mont.) Ignatov & Vanderp. not been reassessed since its description (Ros et al. 2013).

Records. — Bescherelle (1882), Philibert (1889): La Galite, pen-

tes et broussailles du pic central. Feldmann (1947), Jelenc (1955a),
Sérgio & Hébrard (1982): Kabylie de Collo, Baie de Tamanart.
Pterygoneurum subsessile (Brid.) Jur.
Jelenc (1955a): La Calle. Rare.
Pottia chottica Trab.
Pterygoneurum chotticum (Trab.) Broth.
Pseudoscleropodium purum (Hedw.) M.Fleisch.
Nomenclature follows Jelenc (1955a).
Records. — Bescherelle (1882), Jelenc (1955a): Kabylie du Djurd-
jura, Tizi Ouzou. Trabut (1888): sine loco. Feldmann (1947), Jelenc Records. — Battandier & Trabut (1886), Trabut (1887b), Jelenc
(1955a): Grande Kabylie, dans la forêt de Chênes-Lièges de Taourirt (1955a): Prope fontem El Khreider, provinciae Oranensis austra-
Ighil, 900 m; Kabylie de Collo, Bessombour, 500-600 m; Kabylie lis. Trabut (1887a): D’Oran à Mécheria, Chott. Corbière (1904),
de Collo, Taoulel, 500-600 m. Jelenc (1955a): Édough. Rare. Jelenc (1955a): Sétif, sommet d’une coline pierreuse au N-E de la
ville, 1112 m. Brotherus (1924): Algier. Jelenc (1955a): Monts de
Tlemcen, Mazer; Sud algérois, Bou Saâda. Rare.

Pterigynandrum filiforme Hedw.

Ptychostomum capillare
Pterigynandrum filiforme var. majus (De Not.) De Not. (Hedw.) Holyoak & N.Pedersen
Nomenclature follows Hodgetts et al. (2020). Records. — Montagne (1838), Bescherelle (1882), Jelenc (1955a):
Alger. Bescherelle (1882), Lapie (1909): Djurjura. Bescherelle (1882),
Records. — Bescherelle (1882): Djurjura. Bescherelle (1882),
Braun-Blanquet & Maire (1924), Jelenc (1955a): Tlemcen. Bescherelle
Jelenc (1955a): Atlas de Blida, 1400 m; Djebel Babor; Édough;
(1882), Jelenc (1955a): Beni-Salah; La Calle, clairières des bois; Stora;
Kabylie des Babor, Djebel Tababor. Brongniart (1882): Blidah.
Tiaret. Brongniart (1882): Blidah. Corbière (1889): Blida. Corbière
Corbière (1889), Jelenc (1955a): Blida, sommet de l’Abd-el-Kader.
(1904), Jelenc (1955a): Cherchell; Orléansville. Corbière (1905):
Lapie (1909): Kabylie du Djurjura. Jelenc (1953, 1967): Atlas
Environs de Constantine, la Corniche; Environs de Constantine,
de Blida, ravin Bleu de Chréa; Kabylie du Djurdjura, massif de
route de Sétif; Environs de Constantine, Sidi M’Cid; Environs de
l’Aizer, au pic des Cèdres, 1880 m; Massif de l’Aurès, flanc nord
Constantine, tuileries du Rummel. Corbière (1905), Jelenc (1955a,
du djebel Pharaoun, 1800 m; Massif de l’Ouarsenis, cédraie de
1966): Environs de Constantine, djebel Ouach, 1000 m. Jelenc
Téniet el Had, 1500 m. Jelenc (1955a):; Kabylie du Djurdjura,
(1953, 1967): Atlas de Blida, Cèdraies du ravin Bleu à Chrèa; Kabylie
Agoulmin Aberkan; Kabylie du Djurdjura, Aït Ali; Kabylie du
des Babor, flanc nord du djebel Babor, 1700 m; Massif de l’Aurès,
Djurdjura, Akfadou; Kabylie du Djurdjura, Dra Inguel. Jelenc
flanc nord du djebel Pharaoun, 1800 m. Jelenc (1955a): Atlas de
(1955a), Ahayoun et al. (2007): Kabylie du Djurdjura, Tala
Blida, oued el Kébir; Boufarik; Constantine; Corso; Gorges de la
Ghizan. Jelenc (1955a, 1967): Massif de l’Ouarsenis, Téniet el
Chiffa; Kabylie des Babor, Djidjelli; Kabylie du Djurdjura, Azerou
Had, 1500 m. Bizot (1973): Djebel Belezma à 15 km à l’ouest
Tidjer; Kabylie du Djurdjura, Fort National, 1000 m; Kabylie du
de Batna. Frequent.
Djurdjura, Tizi Ouzou; Marengo; Oran; Ouarsenis, djebel Doui;
Ouarsenis, Téniet el Had. Jelenc (1955a, 1967): Alger, La Bouzaréa.
Jelenc (1955a), Ochi (1972): Kabylie des Babor, cap Ras Aokas.
Pterygoneurum ovatum (Hedw.) Dixon Jelenc (1956): Ravin du Frais-Vallon; Sahel d’Alger, environs d’El
Biar. Jelenc (1956, 1967): Massif du Zaccar. Jelenc (1966, 1967):
Records. — Montagne (1838): Alger. Bescherelle (1882): Cons- Kabylie du Djurdjura, massif de l’Akfadou. Jelenc (1967), Ahayoun
tantine; Djebel Mansourah; La Calle, pâturages. Bescherelle (1882), et al. (2007): Massif du Chénoua. Ros et al. (1990): Skikda beach,
Jelenc (1955a): Batna. Trabut (1887a): D‘Oran à Mécheria. Corbière 0 m. Boukhatem et al. (2017): El-Kala National Park, Tonga lake
(1904), Jelenc (1955a): Sétif, sommet d‘une coline pierreuse au N-E watershed. Mazari & Boulaacheb (2023): Setif, Megriss Mountain.
de la ville, 1112 m. Corbière (1905), Jelenc (1955a): Environs de Very common.
Constantine, Chettabah. Braun-Blanquet & Maire (1924), Jelenc
(1955a): Tlemcen. Jelenc (1955a): Berges du Rhumel; La Calle;
Oran; Sud oranais, Kreider. Ros et al. (1990): Ouled Mimoum. Ptychostomum cellulare
Boukhatem et al. (2017): El-Kala National Park, Tonga lake wa- (Hook.) D.Bell & Holyoak
tershed. Frequent.
Bryum cellulare Hook.
Comment
Pottia apoda Durieu ex Besch. nom. nud. was reported by Nomenclature follows Hodgetts et al. (2020).
Bescherelle (1882) based on a sample of Durieu’s herbarium, Records. — Boukhatem et al. (2017): El-Kala National Park,
indicating that it looks like an impoverished form of Ptery- Tonga lake watershed. Very rare.
goneurum cavifolium. Its identity has not been revised since.
Pterygoneurum ovatum var. perraldieri (Besch.) Wijk &
Margad. was described from Algeria as Pottia cavifolia var. Ptychostomum compactum Hornsch.
perraldieri Besch. (Bescherelle 1882). It was reported by Records. — Bescherelle (1882): Kabylie. Jelenc (1955a): Ben Chi-
Bescherelle (1882), Corbière (1905), and Jelenc (1955a) from cao; Kabylie du Djurdjura, Fort National; Kabylie du Djurdjura,
Constantine, Djebel Mansourah, and by Corbière (1905) gorges de l’oued Isser. Rare.

CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5) 93

 Ros R. M. et al.

Ptychostomum donianum Ptychostomum inclinatum (Sw. ex Brid.) J.R.Spence

(Grev.) Holyoak & N.Pedersen
Ptychostomum archangelicum (Bruch & Schimp.) J.R.Spence
Records. — Bescherelle (1882): Birmandreïs, chemins creux; Nomenclature follows Hodgetts et al. (2020) and Holyoak (2021).
Blidah; Constantine, aux moulins; Kabylie; Kaddous; Oran, au
sud du djebel Santo. Bescherelle (1882), Jelenc (1955a): Alger; Records. — Bescherelle (1882), Jelenc (1955a): sine loco.Very rare.
Édough; La Calle; Philippeville; Sidi-Ferruch; Tlemcen, rochers.
Brongniart (1882): Blidah. Gandoger (1883): sine loco. Trabut
(1887a): D’Oran à Mécheria. Cardot (1904), Corbière (1904): Comment
Environs d’Alger, Bouzarea. Corbière (1904), Jelenc (1955a): The only report of the species in Algeria (sub Bryum incli-
Kabylie, Fort National, 1000 m. Litschauer (1904): Jardin de
Marengo in Algier, 50 m. Litschauer (1904), Jelenc (1955a), Ochi natum (Sw. ex Brid.) Turton hom. illeg.) is based on a single
(1972): Frais-Vallon bei la Bouzaréa, 300-400 m. Corbière (1905): specimen collected by Durieu de Maisoneve in Herbarium
Environs de Constantine, Moulin Laire. Camus (1906), Jelenc Motelay without indication of locality (Bescherelle 1882).
(1955a): Oran. Lapie (1909): Kabylie du Djurjura. Jelenc (1949,
1967): Vallée de l’Isser, lieu dit “El Kasbat” près de Lamoricière,
500 m. Jelenc (1955a): Atlas de Blida; Baba Ali; Constantine;
El Biar; Kabylie du Djurdjura, Acherchour n’Tensaout; Kabylie Ptychostomum kunzei (Hornsch.) J.R.Spence
du Djurdjura, Akfadou; Kabylie du Djurdjura, djebel Gouraya;
Khodjaberry; Mascara; Monts de Tlemcen, forêt d’Hafir; Nador Bryum kunzei Hornsch.
de Médéa; Tiaret. Common.
Nomenclature follows Hodgetts et al. (2020).
Records. — Jelenc (1953, 1967): Babor, flanc nord du Babor,
1700 m. Boukhatem et al. (2017): El-Kala National Park, Tonga
Ptychostomum elegans (Nees) D.Bell & Holyoak lake watershed. Rare.
Bryum elegans Nees
Ptychostomum moravicum (Podp.) Ros & Mazimpaka
Nomenclature follows Hodgetts et al. (2020).
Records. — Jelenc (1953, 1967): Massif de l’Aurès, flanc nord du
Records. — Corbière (1905): Environs de Constantine, ravin dejbel Chélia, 2200 m. Jelenc (1967): Ras Kaltoun. Bizot (1973):
de Constantine. Jelenc (1955a): Monts de Constantine, ravin de Djebel Belezma à 15 km à l’ouest de Batna. Rare.
Rhumel. Jelenc (1955a, b, 1967): Tiaret. Rare.

Ptychostomum pallens (Sw.) J.R.Spence

Ptychostomum funkii (Schwägr.) J.R.Spence
Records. — Litschauer (1904): Fort National, 1000 m. Lapie
Bryum funkii Schwägr. (1908, 1909), Trabut (1914), Jelenc (1955a): Kabylie du Djurdjura,
Forêt d’Akfadou. Jelenc (1953, 1967): Ouarsenis, cédraie de Téniet
Nomenclature follows Hodgetts et al. (2020). el Had, 1650 m. Jelenc (1955a): Constantine; Dellys; La Calle.
Jelenc (1966, 1967): Kabylie du Djurdjura, massif de l’Akfadou
Records. — Boukhatem et al. (2017): El-Kala National Park, au Camping Blank, 1500 m. Rare.
Tonga lake watershed. Very rare.

Ptychostomum imbricatulum Ptychostomum pallescens

(Müll. Hal.) Holyoak & N.Pedersen (Schleich. ex Schwägr.) J.R.Spence

Ptychostomum boreale (F.Weber & D.Mohr) Ochyra & Bednarek-

Records. — Montagne (1838), Bescherelle (1882), Jelenc (1955a): Ochyra
Alger. Bescherelle (1882): La Galite. Bescherelle (1882), Jelenc
(1955a): Oran, partie méridionale de la province; Tiaret. Corbière Nomenclature follows Hodgetts et al. (2020).
(1904): Batna, 1600 m. Corbière (1904), Jelenc (1955a): Kab-
ylie, Fort National, 1000 m. Litschauer (1904): Col de Tirourda, Records. — Litschauer (1904): Constantine, auf der Route de la
männlicher Rasen, 1700 m. Corbière (1905), Jelenc (1955a): Corniche, 700 m. Jelenc (1953, 1967): Massif de Ouarsenis, cédraie
Environs de Constantine, Les Pins; Environs de Constantine, de Téniet el Had, 1500 m. Jelenc (1955a): Kabylie du Djurdjura,
ravin du Séminarie; Environs de Constantine, Sidi M’Cid. La- Michelet, Fort National, 1000 m. Rare.
pie (1909): Kabylie du Djurjura. Jelenc (1951, 1967): Massif du
Tessala, pic de Tafaraoui, 650 m. Jelenc (1953): Bèlezma, Tèniet
el Gantos, 1900 m. Jelenc (1953, 1967): Massif de l’Aurès, flanc
nord-ouest du Ras Keltoun, 2200 m. Jelenc (1954b, 1967): Atlas Ptychostomum pseudotriquetrum
Mitidjien, forêt d’El Aioun, 500 m. Jelenc (1955a): Aizer; Arba; (Hedw.) J.R.Spence & H.P.Ramsay
Kreider; La Rassauta; La Réghaia; Mascara; Monts de Batna,
djebel Touggour; Ouarsenis; Tizi-Ouzou. Jelenc (1955a, 1967), Records. — Bescherelle (1882): Alger, El-Biar; La Calle. Be-
Ahayoun et al. (2007): Massif du Bélezma, Téniet el Had. Jelenc scherelle (1882), Jelenc (1955a, 1955b, 1967): Djurjura. Trabut
(1966, 1967): Massif du Bou Zegga, gorges de Keddara, 400 m. (1888): sine loco. Litschauer (1904): Col de Tirourda, 1700 m.
Common. Corbière (1905), Jelenc (1955a): Environs de Constantine, dje-

94 CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5)

 Revisiting the Bryophyte Checklist of Algeria

bel Ouach. Jelenc (1953, 1967): Kabylie du Djurdjura, Aizer, Ptychostomum torquescens
flanc nord du djebel Tachgagalt, 1950 m; Massif de l‘Aurès, flanc (Bruch & Schimp.) Ros & Mazimpaka
nord du djebel Pharaoun. Jelenc (1955a): Alger, La Bouzaréa;
Chenoua; Djebel Dira, Aumale; Gorges de la Chiffa; Guergour. Records. — Bescherelle (1882): Constantine, rochers du Mansourah;
Jelenc (1967): Kabylie du Djurdjura, Aizer, 1750 m; Massif de Djurdjura. Bescherelle (1882), Corbière (1889): Blidah. Bescherelle
l’Aurès, Ain Mimoun. Mazari & Boulaacheb (2023): Setif, Megriss (1882), Jelenc (1955a): Alger; La Calle; Philippeville. Julien (1894):
Mountain. Frequent. Région de Constantine. Corbière (1904), Jelenc (1955a): Cherchell.
Corbière (1905): Environs de Constantine, Bellevue; Environs de
Comment Constantine, Chettabah; Environs de Constantine, djebel Ouach;
Holyoak (2021) confirmed that Bryum pseudotriquetrum var. Environs de Constantine, Les Pins; Environs de Constantine, près
l’usine à gaz; Environs de Constantine, route du Séminarie; Environs
algeriense (Corb.) Wijk & Margad. (synonym: B. ventricosum de Constantine, Sidi M’Cid. Camus (1906), Jelenc (1955a): Oran.
var. algeriense Corb. nom. illeg.), that was reported by Jelenc Jelenc (1949, 1967): Monts de Tlemcen. Jelenc (1951, 1967): Massif
(1955a) from Monts de Constantine, 1000 m, belongs to du Tessala, 730 m; Sahel d’Arzeu, Dejbel (B)Orouse, 630 m. Jelenc
the species Ptychostomum pseudotriquetrum but without (1953): Aurès, flanc nord du djebel Pharaoun, 1800 m. Jelenc (1953,
1967): Massif de l’Ouarsenis, cédraie de Téniet el Had, 1500 m.
assigning to the typical or var. bimum (Schreb.) Holyoak & Jelenc (1954b, 1967): Dune de la Station d’Aquiculture et de Pêche
N.Pedersen. The same applies to B. pseudotriquetrum var. de Castiglione. Jelenc (1955a): Arba; Atlas de Blida, oued el Kebir,
duvalioides Itzigs. in Rabenh. (synonym: Bryum ventricosum 200 m; Azéba; Djebel Chenoua; El Kantara; Frais Vallon, 300 m;
var. duvalioides (Itzigs.) Glow.) and Bryum pseudotriquetrum Kabylie des Babor, cap Aokas; Kabylie du Djurdjura, Aït Khalfoun;
var. gracilescens Schimp. (synonym: Bryum ventricosum var. Kabylie du Djurdjura, Akfadou; Kabylie du Djurdjura, djebel Gouraya;
La Bouzaréa; Lambèse; Monts de Constantine, gorges de Rhumel;
gracilescens (Schimp.) Lindb. nom. inval.). djebel Murdjadjo; Ouarsenis. Jelenc (1955a, 1967): Saïda. Jelenc
(1956, 1967): Kabylie du Djurdjura, près de Tizi-Ouzou; Massif
de la Bouzaréa, forêt de Baïnem; Massif de la Bouzaréa, ravin près
Ptychostomum rubens des Bains Romains. Bizot (1973): Ruines de Timgad. Syed (1973):
(Mitt.) Holyoak & N.Pedersen Kabylia minor, Ras Aokas. Ros et al. (1990): Skikda beach, 0 m.
Ahayoun et al. (2007): sine loco. Mazari & Boulaacheb (2023): Setif,
Megriss Mountain. Very common.
Bryum trabutii Thér.
Nomenclature follows Ros et al. (2023).
Comment
Records. — Thériot & Trabut (1930), Jelenc (1955a), Ros et al. Ros et al. (1990) reported Ptychostomum gr. torquescens from
(2023): Numidia prope La Calle. Very rare. Oued Rhiou.

Ptychostomum schleicheri
(DC.) J.R.Spence ex D.Bell & Holyoak Ptychostomum turbinatum (Hedw.) J.R.Spence

Bryum schleicheri DC. Bryum turbinatum (Hedw.) Turner

Bryum turbinatum var. gracilifolium Besch.
Bryum schleicheri var. latifolium (Schwägr.) Schimp.
Nomenclature follows Hodgetts et al. (2020) and Holyoak (2021).
Ptychostomum schleicheri var. latifolium (Schwägr.) D.Bell & Holyoak
Records. — Bescherelle (1882): Atlas; Constantine, chute du Rum-
Nomenclature follows Holyoak (2021). mel; La Calle, pentes gazonnées humides. Lapie (1909): Kabylie du
Records. — Corbière (1904), Jelenc (1955a): Bône, ravin aux Djurjura. Jelenc (1951, 1967): Haute plaine de Tlemcen, cascades
flancs de l’Édough, 200 m. Trabut (1914), Jelenc (1955a): Djurd- de l’oued bou Ennag à la traversée des collines des Beni Mester,
jura. Thériot (1931, 1933): Atakor-n-Ahaggar; Tarouda, 2100 m; 500 m, 650 m. Jelenc (1955a): Blida. Jelenc (1955a, 1956, 1967):
Tehi-n-Tekart, 2000 m. Thériot (1931, 1933), Jelenc (1955a): Ahag- Kabylie du Djurdjura, Main du Juif, 1600 m. Rare.
gar, Ideles, 1500 m; Hoggar, Imarera, 2000 m; Hoggar, Tessekimt,
1900 m; Tamanghasset, 1500 m; Tefedest, Ahetes, 1150 m. Thériot
(1931), Jelenc (1955a), Frahm (1978): Hoggar, Gueltas von Is- Pulvigera lyellii
sakarassene, 1880 m. Frahm (1978): Hoggar, Gueltas von Afilale, (Hook. & Taylor) Plášek, Sawicki & Ochyra
2000 m. Frequent.
Orthotrichum lyellii Hook. & Taylor

Comment Nomenclature follows Plášek et al. (2015).

Bryum schleicheri fo. peracutifolium Thér. and B. schleicheri fo. Records. — Bescherelle (1882): Atlas, troncs d’arbres; Bouzazi.
stenophyllum were described from Algeria by Thériot (1931) Bescherelle (1882), Jelenc (1955a): Kabylie du Djurdjura, Tizi Ou-
from Hoggar, Tamanghasset (in Tehi- n-Tekart, at 2000 m zou; La Calle. Corbière (1889): Blida, forêt des Beni-Salah, 1500 m.
the first, and in Tessekimt at 1900 m the second), and later Corbière (1905): Environs de Constantine, djebel Ouach. Lapie
(1909): Kabylie du Djurdjura, Beni Khalfoun. Jelenc (1949, 1967):
compiled in Jelenc (1955a). Its identity has not been reas- Monts de Tlemcen, forêts de Zariflet et d’Hafir, 1000-1400 m.
sessed since its description (Ros et al. 2013). They are probably Jelenc (1955a): Kabylie de Collo, djebel Goufi; Kabylie du Djurd-
synonyms of P. schleicheri. jura, Aït Daoud; Monts de Constantine; Téniet el Had. Ahayoun

CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5) 95

 Ros R. M. et al.

et al. (2007): Djebel Bonti, Collo. Boukhatem et al. (2017): El-Kala près de Bou Touya, 600 m. Jelenc (1954b, 1956, 1967): Kabylie
National Park, Tonga lake watershed. Mazari & Boulaacheb (2023): du Djurdjura, massif de l’Akfadou, à la Fontaine des Ifs, 1200 m.
Setif, Megriss Mountain. Frequent. Jelenc (1955a): Djidjelli. Boukhatem et al. (2017): El-Kala National
Park, Tonga lake watershed. Rare.

*Pylaisia polyantha (Hedw.) Schimp. Rhynchostegiella curviseta (Brid.) Limpr.

Records. — Trabut (1887a): D’Oran à Mécheria. Hypnum algirianum (Brid. ex P.Beauv.) Brid.
Rhynchostegiella algiriana (Brid. ex P.Beauv.) Warnst.
Comment
Doubtfully present in Algeria. The report of this Circum- Nomenclature follows Patiño et al. (2017).
polar Temperate species (Blockeel et al. 2014b) in Algeria Records. — Desfontaines (1799), Gandoger (1883): sine loco.
is based on a single specimen, and its reliability is doubtful. Lamarck (1813), Montagne (1838), Bescherelle (1882), Jelenc
Due to the difficulty in locating this collection to be revised, (1955a), Ahayoun et al. (2007): Environs d’Alger. Bescherelle
the species can only remain in the Algerian catalogue if new (1882), Corbière (1904), Jelenc (1955a): Bône. Bescherelle (1882),
specimens are found. Jelenc (1955a): Blidah; La Calle; Tlemcen. Corbière (1904), Jelenc
(1955a): La Bouzaréah, près Alger. Jelenc (1949, 1967): Massif du
Pylaisia polyantha fo. gracillima Besch. was described by Murdjadjo (Oran), Forêt de Msila, ravin de la source de Guedda-
Bescherelle (1882) from Sahara d’Oran, entre Chellala et ra, 180 m. Jelenc (1951, 1967): Massif du Tessala, pic Tafaraoui,
Arba-Tatani, and later compiled in Jelenc (1955a). According 600 m. Jelenc (1953, 1967): Massif de l’Aurès, forêt de djebel Sgag,
to Ros et al. (2013), its identity has not been revised since. 1700 m. Jelenc (1955a): Birkadem; Djebel Beni Smir; Edough;
Its taxonomic value is doubtful. Gouraya. Jelenc (1956, 1967), Jovet-Ast (1956): Atlas Mitidjien, pic
Mouzaïa. Boukhatem et al. (2017): El-Kala National Park, Tonga
lake watershed. Frequent.

Pyramidula tetragona (Brid.) Brid. Comment

The criterium of Patiño et al. (2017) has been followed
Records. — Douin (1904): Aux environs de Constantine, 1000 m.
Corbière (1905), Jelenc (1955a): Monts de Constantine, djebel for considering the reports of the genus Rhynchostegiella
Ouach. Rare. in Algeria. These authors excluded R. letourneuxii of the
synonyms of R. curviseta and included it within Rhyn-
chostegium confertum. Also, they considered Rhynchostegiella
Racomitrium aciculare (Hedw.) Brid. algiriana a synonym of R. curviseta, instead of R. tenella as
was considered in Ros et al. (1999, 2013), and in Jelenc
Records. — Jelenc (1948), Bednarek-Ochyra (2006): Kabylie, Ba-
bor. Jelenc (1955a), Bednarek-Ochyra (2006): Kabylie du Djurdjura, (1955a). Therefore, the reports of Rhynchostegiella curviseta
Aït Idjer; Kabylie du Djurdjura, Tala Ghizan; Kabylie du Djurdjura, here included are: 1) the original reports under the names
Tamesguida; Kef Mcid el Aïssa, Dépt. de Constantine. Rare. Rhynchostegiella curviseta (Brid.) Limpr. and Rhynchostegium
curvisetum (Brid.) Schimp., and 2) the reports of Rhychost-
egiella algiriana and their homotypic synonyms (Hypnum
Racomitrium affine (F.Weber & D.Mohr) Lindb. algirianum (Brid. ex P.Beauv.) Brid. and Rhynchostegium
algirianum (Brid. ex P.Beauv.) Lindb.) but excluding those
Records. — Corbière (1905): Environs de Constantine, djebel of Jelenc (1955a, 1956, 1967), whose identity is doubtful
Ouach. Boukhatem et al. (2017): El-Kala National Park, Tonga
lake watershed. Rare. as can correspond to any of both species [Jelenc (1955a):
Constantine; Frais Vallon; Kabylie des Babor, Cap Aokas;
Kabylie du Djurdjura, Akfadou; Kabylie du Djurdjura,
Racomitrium canescens (Hedw.) Brid. Yakouren. Jelenc (1956): Massif de la Bouzaréa, ravin de la
prise d’eau près des Bains-Romains. Jelenc (1956, 1967):
Records. — Bescherelle (1882), Jelenc (1955a): Kabylie du Djurd- Sahel d’Alger, environs d’El Biar. Jelenc (1967): Massif du
jura, Tizi Ouzou. Very rare.
Bou Zegza, gorges de Keddara, 400 m].
Given the great taxonomical changes proposed in the last
Racomitrium heterostichum (Hedw.) Brid. decades, the identity of old Algerian Rhynchostegiella reports
is unsure and therefore, the presence and distribution of the
Records. — Corbière (1905), Jelenc (1955a): Monts de Constan- species in the country should be reevaluated.
tine, djebel Ouach. Very rare.

Rhizomnium punctatum (Hedw.) T.J.Kop. Rhynchostegiella litorea (De Not.) Limpr.

Records. — Feldmann (1947), Jelenc (1955a): Kabylie de Collo, Rhynchostegiella tenella var. meridionalis (Boulay) Zodda
l’oued Medjerrah près de Bou Noghra, 700 m; Kabylie de Collo,
près de Taoulel, 600 m; Kabylie de Collo, source à Krerach Mayoun Nomenclature follows Patiño et al. (2017).

96 CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5)

 Revisiting the Bryophyte Checklist of Algeria

Records. — Bescherelle (1882): Mustapha. Bescherelle (1882), Rhynchostegium confertum (Dicks.) Schimp.
Jelenc (1955a): Alger; Bouzaréah; Staouëli. Corbière (1904):
Telemly près Alger. Corbière (1904), Jelenc (1955a): Gorges de Rhynchostegiella letourneuxii (Besch.) Broth.
la Chiffa, ruisseau des Singe. Dixon (1904): in the neighbour-
hood of Algiers. Litschauer (1904), Jelenc (1955a): Vallon bei Rhynchostegium letourneuxii Besch.
la Bouzarea, 300 m. Corbière (1905): Environs de Constantine,
Moulin Laire. Corbière (1905), Jelenc (1955a): Environs de Nomenclature follows Patiño et al. (2017).
Constantine, bords de l’oued Bou Merzoug, 550 m. Feldmann
(1947), Jelenc (1955a): Kabylie de Collo, les crêtes de Boudra,
Records. — Bescherelle (1882), Jelenc (1955a): Djebel Édough,
500-600 m. Jelenc (1951): Monts de Tlemcen, Koudiat d’Hafir,
ravin de Ringa et sur le sentier du bas; Djebel Édough, sentier
1250 m. Jelenc (1954b): Sahel d’Alger, Bois de Boulogne. Jelenc
du milieu; La Calle, marais du Tonga; La Chiffa; Oued El-Aneb.
(1954b, 1967): Massif du Bou Zegza, gorges de Keddara, 400 m.
Corbière (1889): Blida, derrière Mimich. Corbière (1904), Jelenc
Jelenc (1955a): Atlas de Blida; Birkadem; Constantine. Jelenc
(1955a): Gorges de la Chiffa, ruisseau des Singes. Corbière
(1956, 1967): Massif du Bou Zegza, dans les gorges de Keddara,
(1904, 1905), Jelenc (1955a): Environs de Constantine, Route
400 m; Sahel d‘Alger, colline du Jardin d‘Essai. Jelenc (1967):
de la Corniche, les Pins, Dj. Ouach, 1000 m, 850 m. Litschau-
Monts de Tlemcen, Koudiat d‘Hafir, 1250 m. Düll (1986): sine
er (1904), Jelenc (1955a): Bône, 50 m. Camus (1906): Oran.
loco. Boukhatem et al. (2017): El-Kala National Park, Tonga lake
Braun-Blanquet & Maire (1924): Tlemcen. Jelenc (1955a): Kaby-
watershed. Common.
lie des Babor, cap Aokas; Kabylie du Djurdjura, djebel Afroun;
Kabylie du Djurdjura, forêt d’Akfadou; Kabylie du Djurdjura,
Comment Tala Ghizan; La Bouzaréa; Monts de Tlemcen. Jelenc (1967):
sine loco. Frequent.
The reports included here are the original reports those of
Rhynchostegiella litorea, and their homotypic synonyms
(Rhynchostegiella tenella var. litorea (De Not.) P.W.Richards & Comment
E.C.Wallace, and Rhynchostegium litoreum (De Not.) Bott.), Rhynchostegium confertum var. latifolium Renauld in Jelenc
as well as those of Rhynchostegiella tenella var. meridionalis nom. inval. was reported by Jelenc (1955a) from Monts de
and their homotypic synonyms (Rhynchostegiella algiriana Constantine, djebel Ouach, but according to Ros et al. (2013),
var. meridionalis (Boulay) Corb., Rhynchostegium algirianum its identity has not been revised since.
var. meridionale (Boulay) Corb. nom. illeg., and Rhynchoste-
gium tenellum var. meridionale (Boulay) Paris).
Rhynchostegium megapolitanum
(Blandow ex F.Weber & D.Mohr) Schimp.
Rhynchostegiella tenella (Dicks.) Limpr.
Records. — Bescherelle (1882): Birmandreïs; Kabylie; Mus-
Records. — Bescherelle (1882): Alger; Birkadem; Édough, près tapha, ravins de l’Agha. Bescherelle (1882), Jelenc (1955a): Al-
la route Randon; Bougie; Oran; Philippeville; Tlemcen. Trabut ger; Bône; Dunes des Beni-Urgin; La Calle; Vallée de Biskader.
(1887a): D’Oran à Mécheria. Julien (1894): Région de Constan- Gandoger (1883), Trabut (1888): sine loco. Corbière (1889):
tine. Ros et al. (1990): Djebel Chenoua, 5-40 m. Boukhatem et al. Blida. Corbière (1904): Alger, le Bois de Boulogne. Corbière
(2017): El-Kala National Park, Tonga lake watershed. Frequent. (1904), Jelenc (1955a): Médéa; Orléansville, bois de la Pépin-
ière. Litschauer (1904), Jelenc (1955a): Chiffaschlucht und im
Tal des Qued-Kebir bei Blidah, 300 m. Corbière (1905), Jelenc
Comment (1955a): Environs de Constantine, Bellevue; Environs de Con-
stantine, djebel Ouach; Environs de Constantine, Le Hamma;
Only the original Algerian reports sub Rhynchostegiella tenella Environs de Constantine, Les Pins; Environs de Constantine,
(Dicks.) Limpr. and Rhynchostegium tenellum (Dicks.) Schimp. Menidj; Environs de Constantine, pont d’Aumale; Environs de
are included here. Constantine, près les Arcades romaines. Braun-Blanquet & Maire
(1924), Jelenc (1955a): Tlemcen. Jelenc (1949, 1967): Monts
de Tlemcen, djebel el Béniane, 950 m. Jelenc (1951, 1967):
Massif du Tessala, pic Tafaraoui, 550 m. Jelenc (1953, 1967):
Rhynchostegiella teneriffae (Mont.) Dirkse & Bouman Kabylie du Djurdjura, massif de l’Aizer, flanc nord du djebel
Tachgagalt, 2000 m. Jelenc (1954b, 1967): Chaîne des Zahrez,
Records. — Bescherelle (1882), Jelenc (1955a): La Calle. Corbière djebel Touilila, 1000 m, l’oued M’Kamza. Jelenc (1955a): Alger,
(1889): Blida, dans un ruisselet derrière Mimich. Jelenc (1955a): Tixérain; Atlas de Blida; Kabylie des Babor, cap Aokas; Kabylie
Atlas de Blida; La Bouzaréa. Rare. du Djurdjura, Tirourda; La Bouzaréa; Maison-Carrée. Jelenc
(1956, 1967): Massif de la Bouzaréa, forêt de Baïnem; Massif
du Dahra, Petit-Port dans la callitriaie. Ros et al. (1990): Ham-
Comment mam-Riga, 500 m. Mazari & Boulaacheb (2023): Setif, Megriss
Mountain. Common.
According to Dirkse & Bouman (1995), the presence of this
species in Algeria is based on the reports of Eurhynchium
teesdalei (Schimp.) Milde and Rhynchostegiella teesdalei Comment
(Schimp.) Limpr. Although Patiño et al. (2017) considered Rhynchostegium megapolitanum var. hygrophyllum Renauld
this synonymy doubtful, Hodgetts et al. (2020) maintai- in Jelenc nom. inval. was reported by Jelenc (1955a) from
ned it pending further work, and this is the criterium also Monts de Constantine, djebel Ouach, but according to Ros
followed here. et al. (2013), its identity has not been revised since.

CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5) 97

 Ros R. M. et al.

Rhynchostegium murale (Hedw.) Schimp. Djurdjura, Tablabalt; Monts de Tlemcen, Mazer; Ouarsenis, Té-
niet el Had. Jelenc (1956): Monts de Tlemcen, vallée de l’oued
Records. — Montagne (1838): Alger. Bescherelle (1882), Jelenc Isser près de Lamoricière. Jelenc (1967): Vallée de l’oued Isser à
(1955a): Édough; Gorges de la Chiffa; Tlemcen. Trabut (1888): Lamoricière. Frequent.
sine loco. Jelenc (1955a): Bône; Kabylie du Djurdjura, Agoulmin
Aberkan; Kabylie du Djurdjura, Akfadou. Rare.
Comment
As stated by Ros et al. (2013), the reports of Schistidium
Rhynchostegium riparioides (Hedw.) Cardot apocarpum complex in the Mediterranean published before
the revision of Blom (1996) should be revised to establish
Records. — Bescherelle (1882): Blidah; Djurjura; Oued El-Af- to which of the presently accepted species they correspond.
ran. Bescherelle (1882), Jelenc (1955a): Édough; Oran. Trabut
(1887a): D’Oran à Mécheria. Corbière (1889): Blida, derrière
Mimich. Corbière (1904), Jelenc (1955a): Tlemcen, parois de la
cascade. Litschauer (1904), Jelenc (1955a): Kabylie du Djurdjura, Schistidium confertum (Funck) Bruch & Schimp.
Fort National, 1000 m. Corbière (1905): Environs de Constan-
tine, djebel Ouach, 900 m. Lapie (1909): Kabylie du Djurjura. Records. — Cardot (1904), Maire (1916), Jelenc (1955a):
Feldmann (1947), Jelenc (1955a): Kabylie de Collo, oued Bin el Province d’Oran, Djebel Morghad, 1950 m, 2105 m. Corbière
Ouidane près de Taoulel. Jelenc (1949, 1967): Haute plaine de (1905), Jelenc (1955a): Environs de Constantine, Sidi M’Cid.
Tlemcen, oued Saf-Saf entre le village et le cirque d’el Ourit; Monts Jelenc (1956, 1967): Massif du Bou Zegza, dans les gorges de
de Tlemcen, forêt de Zariffet aux cascades de l’oued Ouadallah. Keddara, 400 m. Rare.
Jelenc (1951): Monts de Haute plaine de Tlemcen. Jelenc (1953,
1967): Babor, flanc nord du djebel Babor, 1300 m; Massif de
l’Aurès, forêt de Sgag, 1650 m; Massif de l’Aurès, maison forestière
du djebel Chélia, 1500 m. Jelenc (1955a): Atlas de Blida; Aurès, Schistidium crassipilum H.H.Blom
djebel Chélia, 1350 m; Djebel Dira; Gorges de la Chiffa; Kabylie
du Djurdjura, Akfadou; Kabylie du Djurdjura, Ansor Aberkane; Records. — Bescherelle (1882), Jelenc (1955a), Ros & Cano (2008):
Kabylie du Djurdjura, Dra Inguel; Kabylie du Djurdjura, Tiro- Aurès. Jelenc (1955a): Lambèse; Ouarsenis, Téniet el Had. Boukhatem
urda; Koléa; Maison-Carrée. Jelenc (1967): Kabylie des Babor, et al. (2017): El-Kala National Park, Tonga lake watershed. Rare.
Tamerdjajout. Boukhatem et al. (2017): El-Kala National Park,
Tonga lake watershed. Common.
Schistidium flaccidum (De Not.) Ochyra

Comment Records. — Cardot (1904), Maire (1916), Jelenc (1955a): Pro-

Platyhypnidium rusciforme var. squarrosum (Boulay) Latzel vince d’Oran, Djebel Morghad, 1950 m, 2015 m. Jelenc (1955a):
Atlas de Blida, oued el Kebir, 200 m; Kabylie du Djurdjura, col de
was reported by Jelenc (1955a) from Monts de Constantine, Tirourda, 1700 m; Kabylie du Djurdjura, Fort National; Kabylie
djebel Ouach, but according to Ros et al. (2013), its identity du Djurdjura, Michelet, 1000-1200 m. Bremer (1980): Oran.
has not been revised since. Also, it is the case of two invalid Boukhatem et al. (2017): El-Kala National Park, Tonga lake wa-
names: Platyhypnidium rusciforme var. longifolium Trab. in tershed. Rare.
Jelenc nom. inval. descr. gall., proposed by Jelenc (1955a)
from Djebel Dira and Kabylie du Djurdjura, Tirourda, and
Platyhypnidium rusciforme fo. tophacea Besch. nom. nud., pro- *Sciuro-hypnum plumosum
posed by Bescherelle (1882) without precise locality in Algeria (Hedw.) Ignatov & Huttunen
(sine loco), based on a specimen of Durieu de Maisonneuve
and later compiled by Jelenc (1955a). Records. — Bescherelle (1882), Jelenc (1948, 1955a): Kabylie,
Tala-Guizan. Jelenc (1955a): Kabylie du Djurdjura, Akfadou.

Schistidium agassizii Sull. & Lesq. Comment

Records. — Jelenc (1953, 1967): Kabylie du Djurdjura, Aizer, flanc
Doubtfully present in Algeria. The presence of this Circumpo-
nord du djebel Tachgagalt, 1950 m. Jelenc (1955a): Ouarsenis. Jelenc lar Boreo-temperate species (Blockeel et al. 2014b) in Algeria
(1956, 1967): Atlas Mitidjien, pic Mouzaïa, 1200 m. Boukhatem is unlikely. Given the challenges in locating the collections,
et al. (2017): El-Kala National Park, Tonga lake watershed. Rare. this species could only be retained in the Algerian catalogue
upon the discovery of new specimens. Orgaz et al. (2011) did
not confirmed any north African report.
Schistidium apocarpum (Hedw.) Bruch & Schimp.

Records. — Bescherelle (1882): Djurdjura; Petit Atlas. Besche-

relle (1882), Jelenc (1955a): Constantine; Djebel Babor. Corbière Sciuro-hypnum populeum
(1904), Jelenc (1955a): Sétif, 1100 m. Corbière (1905): Environs (Hedw.) Ignatov & Huttunen
de Constantine, La Corniche. Jelenc (1949, 1967): Monts de
Tlemcen, pentes du djebel el Béniane, 850-1000 m. Jelenc (1951, Records. — Bescherelle (1882), Jelenc (1955a): Aurès; Djurjura.
1967): Massif du Tessala, pic de Tafaraoui, 650 m. Jelenc (1955a): Jelenc (1955a): Babor; Kabylie du Djurdjura, Akfadou; Kabylie du
Alger; Atlas de Blida, oued el Kebir, 200 m; Aurès; Kabylie du Djurdjura, Tala Ghizan. Orgaz et al. (2011): sine loco. Rare.

98 CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5)

 Revisiting the Bryophyte Checklist of Algeria

Scleropodium touretii (Brid.) L.F.Koch (1966): Monts de Tlemcen, forêt de Zariflet, 1150 m. Ros et al.
(1990): Between Mostaganem and Cap Ivi, Abdelmaimalek-Ram-
Records. — Montagne (1838), Bescherelle (1882), Jelenc (1955a): dan beach, 0 m; Djebel Chenoua, 5-40 m. Ahayoun et al. (2007):
Alger. Bescherelle (1882): Kabylie. Bescherelle (1882), Lapie (1909): Oran. Boukhatem et al. (2017): El-Kala National Park, Tonga lake
Djurjura. Bescherelle (1882), Jelenc (1955a): Bône; Constantine; La watershed. Very common.
Calle; Philippeville; Tiaret. Gandoger (1883), Trabut (1888): sine
loco. Corbière (1889): Blida. Corbière (1904): La Bouzaréah, près
Alger. Corbière (1904, 1905): Environs de Constantine, djebel Ou- Scorpiurium deflexifolium
asch, 1100 m. Corbière (1904), Litschauer (1904), Jelenc (1955a): (Solms) M.Fleisch. & Loeske
Kabylie, Fort National, Michelet, 1000-1200 m. Corbière (1904),
Jelenc (1955a): Nador de Médéa. Litschauer (1904): Philippeville, Records. — Corbière (1904), Jelenc (1955a): Bône, ruisseau aux
50 m. Litschauer (1904), Jelenc (1955a): Kabylie du Djurdjura, Col flancs de l’Édough. Corbière (1905), Jelenc (1955a): Environs de
de Tirourda, 1700 m; Vallon bei Alger, 300 m. Litschauer (1904), Constantine, bords de l’oued Bou Merzoug, 550 m. Jelenc (1951,
Jelenc (1955a), Ros et al. (1990): Chiffaschlucht und im Tal des 1967): Monts de Tlemcen, forêt de Zariflet, 1150 m. Jelenc (1955a):
Qued-Kebir bei Blidah, 200-300 m. Corbière (1905): Environs de La Calle; Stora. Boukhatem et al. (2017): El-Kala National Park,
Constantine, ravin du Séminarie. Corbière (1905), Jelenc (1955a): Tonga lake watershed. Rare.
Environs de Constantine, Azeba. Feldmann (1947), Jelenc (1955a):
Kabylie de Collo, Baie de Tamanart. Jelenc (1949, 1967): Monts
de Tlemcen, pentes du djebel el Béniane, 1000 m, dans la “Forêt
des Pins”. Jelenc (1951, 1967): Sahel d’Oran, plateau des Ghamras, **Sematophyllum demissum (Wilson) Mitt.
400 m. Jelenc (1953, 1967): Atlas de Blida, ravin Bleu de Chréa;
Massif de l’Aurès, flanc nord du djebel Pharaoun, 1800 m. Jelenc Comment
(1955a): Kabylie des Babor, cap Aokas; Kabylie du Djurdjura, Aït
Khalfoun; Kabylie du Djurdjura, Tala Ghizan; La Réghaia. Jelenc Excluded from Algeria. The only report was made by Feld-
(1956): Sahel d’Alger, colline du Jardin d’Essai. Jelenc (1956, 1967): mann (1945, 1947) and compiled by Jelenc (1955a) from
Atlas Mitidjien, Meurdja, 1200 m; Massif du Tessala, pic Tessala, Kabylie de Collo, Oued Elli el Diab. Pierrot (2000) revised
1000 m; Massif du Zaccar. Jelenc (1966): Monts de Tlemcen, the specimen kept in the Faurel herbarium and concluded it
forêt de Zariffet, 1100 m. Jelenc (1967): Atlas Mitidjien, forêt d’El corresponds to S. substrumulosum.
Aioun, 500 m; Atlas Mitidjien, pic Mouzaïa; Kabylie du Djurdjura,
massif de l’Akfadou au Camping Blank. Ros et al. (1990): Djebel
Chenoua, 5-40 m; Skikda beach, 0 m; Texenna’s wood, 700 m.
Mazari & Boulaacheb (2023): Setif, Megriss Mountain. Common. Sematophyllum substrumulosum (Hampe) E.Britton
Records. — Bescherelle (1882): Kabylie; Ras-Aokas. Feldmann
(1945, 1947), Jelenc (1955a), Pierrot (2000): Kabylie de Collo,
Scorpiurium circinatum (Bruch) M.Fleisch. & Loeske oued Elli el Diab, près du Melhab. Feldmann (1945), Jelenc (1955a):
Près d’Alger, fôret de Baïnem. Rare.
Thamnium alopecurum var. decipiens Corb.

Nomenclature follows Ros et al. (2023). Sphagnum auriculatum Schimp.

Records. — Bescherelle (1882): Kabylie; La Calle, sur les pierres Sphagnum subsecundum var. algerianum Cardot
inondées; Stora. Bescherelle (1882), Corbière (1904), Litschauer
(1904), Jelenc (1955a): Alger. Bescherelle (1882), Jelenc (1955a): Nomenclature follows Eddy (1985).
Bône; Constantine; Philippeville. Gandoger (1883), Trabut (1888):
sine loco. Corbière (1889): Blida. Corbière (1904): Tlemcen, ro- Records. — Bescherelle (1882), Cardot (1884), Jelenc (1955a):
chers entre Sidi-Bou-Médine. Corbière (1904), Jelenc (1955a): Bône, marais dans le Senhadja; La Calle, petit marais au sud, sur le
Médéa. Litschauer (1904): Constantine auf der Route de la Cor- flanc d´une colline boisée and bruyères marécaugeuses. Bescherelle
niche, 700 m. Litschauer (1904), Jelenc (1955a): Vallon bei la (1882), Jelenc (1955a): La Calle, petit marais au sud, sur le flanc
Bouzarea, 400 m. Litschauer (1904), Jelenc (1955a), Ros et al. d’une colline boisée. Gauthier-Lièvre (1931): 24 km from El Kala.
(1990): Chiffaschlucht und im Tal des Qued-Kebir bei Blidah, Gauthier-Lièvre (1931), Jelenc (1967): Marais de Bou Merchen
200-300 m. Corbière (1905): Environs de Constantine, Les betwenn Lac Melah and El Kala. Jelenc (1955a): Plaine de Bône,
Pins; Environs de Constantine, ravin de Constantine; Environs vallé de l’oued Aouicha entre le Bou Zioun et le Kef Zouara. Muller
de Constantine, Sidi M’Cid. Camus (1906), Braun-Blanquet & et al. (2010): Aïn Bergougaïa. Boukhatem et al. (2017): El-Kala
Maire (1924), Jelenc (1955a): Tlemcen. Lapie (1909): Kabylie du National Park, Tonga lake watershed. Rare.
Djurjura. Feldmann (1947), Jelenc (1955a): Kabylie de Collo, Baie
de Tamanart. Jelenc (1949, 1967): Massif du Murdjadjo (Oran),
ravin de l’oued Tammermouth. Jelenc (1951, 1967): Haute-Plaine **Sphagnum contortum Schultz
de Tlemcen, cascades de l’oued bou Ennag à la traversée des col-
lines des Beni-Mester, 500 m; Massif du Tessala, pic de Tafaraoui,
700 m; Monts de Tlemcen, forêt de Tessera Mramet, 1400 m. Comment
Jelenc (1954b, 1967): Atlas Mitidjien, forêt d’El Aioun, 500 m; Excluded from Algeria. It was reported as S. subsecundum var.
Massif de l’Akfadou, Camping Black, 1500 m; Massif du Bou contortum (Schultz) Huebener by Bescherelle (1882) from La
Zezga, gorges de Keddara, 400 m. Jelenc (1955a): Alma; Cherchell; Calle, petit marais au sud, sur le flanc d´une colline boisée
Djurdjura; Gorges de la Chiffa, ruisseau des Singes; Kabylie des
Babor, cap Aokas; Monts de Tlemcen, forêt d’Hafir; Tiaret, oued and by Gauthier-Lièvre (1931) from Marais de Bou Merchen
Mina. Jelenc (1955a, 1967): Sahel d’Alger, massif de la Bouzaréa. betwenn Lac Melah and El Kala. According to Muller et al.
Jelenc (1956): Sahel d’Alger, au-dessus du Jardin d’Essai. Jelenc (2010), only S. auriculatum is present in Algeria.

CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5) 99

 Ros R. M. et al.

**Sphagnum palustre L. Syntrichia laevipila Brid.

Comment Records. — Bescherelle (1882): Kabylie; Petit Atlas. Bescherelle

(1882), Corbière (1889), Gallego (2005): Blidah. Bescherelle (1882),
Excluded from Algeria. It was reported as S. cymbifolium (Ehrh.) Jelenc (1955a): Bône; Édough; La Calle; Philippeville. Bescherelle
Hedw. by Jelenc (1955a) from Plaine de Bône, vallé de l’oued (1882), Jelenc (1955a), Gallego (2005): Bouzaréah. Bescherelle
Aouicha entre le Bou Zioun et le Kef Zouara. According to Muller (1882), Jelenc (1955a), Gallego (2005), Ahayoun et al. (2007):
et al. (2010), only Sphagnum auriculatum is present in Algeria. Alger. Gandoger (1883), Barkman (1963): sine loco. Trabut (1887a):
D’Oran à Mécheria, Tell. Corbière (1904): Tlemcen, à Sidi-Bou-
-Médine. Corbière (1904), Jelenc (1955a): Médéa; Sétif, 1080 m.
Corbière (1905): Environs de Constantine, sur les arbres du square.
**Sphagnum subsecundum Nees Lapie (1909): Kabylie du Djurjura. Jelenc (1951, 1967): Monts de
Tlemcen, forêt de Zariffet près de l’aïn Defla. Jelenc (1955a): Aïn
Comment Taya; Alma; Atlas de Blida; Constantine; Frais Vallon, 300 m; Kabylie
des Babor, cap Aokas; Kabylie des Babor, djebel Tababor; Kabylie
Excluded from Algeria. It was reported as Sphagnum subsecun- du Djurdjura, Akfadou; Kabylie du Djurdjura, djebel Gouraya;
dum and its synonym S. laricinum (Wilson) Spruce ex Lindb. Kabylie du Djurdjura, Fort National; Khodjaberry; Mascara; Mou-
by Bescherelle (1882) and Jelenc (1955a) from Bône, marais zaïaville; Tlemcen; Vallée du Chéliff, Duperré. Jelenc (1966): Monts
dans le Souhadja and from La Calle, bruyères marécaugeuses, de Constantine au djebel Ouach. Jelenc (1967): Kabylie de Collo;
and by Gauthier-Lièvre (1931) and Jelenc (1967) from Marais Massif du Bou Zegza, gorges de Keddara, 400 m. Gauthier (1987):
Assekrem, Hoggar. Gallego (2005): Tizi-Ouzou. Boukhatem et al.
de Bou Merchen between Lac Melah and El Kala. Accord- (2017): El-Kala National Park, Tonga lake watershed. Mazari &
ing to Muller et al. (2010), only Sphagnum auriculatum is Boulaacheb (2023): Setif, Megriss Mountain. Common.
present in Algeria.

Syntrichia montana Nees

Streblotrichum convolutum (Hedw.) P.Beauv.
Records. — Bescherelle (1882): Sommet de l’Atlas central. Bescherelle
Barbula convoluta Hedw. (1882), Jelenc (1955a): Constantine, rochers du ravin. Cardot (1904),
Maire (1916), Jelenc (1955a): Province d’Oran, Djebel Morghad,
Nomenclature follows Kučera et al. (2013). 1950 m, 2105 m. Corbière (1905): Environs de Constantine, Les
Pins. Jelenc (1949, 1967): Monts de Tlemcen, pentes au sud de Tle-
Records. — Bescherelle (1882): Kabylie. Bescherelle (1882), Jelenc mcen, 850-1000 m. Jelenc (1951), Jelenc (1967): Massif du Tessala,
(1955a): Alger; Bône; Bouzaréah; Tiaret. Corbière (1904), Jelenc pic de Tafaraoui, 650 m. Jelenc (1953, 1967): Massif du Bélezma,
(1955a): Orléansville, bois de la Pépinière. Jelenc (1955a): Akfadou; cédraies de l’oued Chaba, 1400 m. Jelenc (1955a): Atlas de Blida,
Biban, Constantine; Cap Aokas; Chebka, oued Mredjar au Nord oued el Kebir, 200 m. Aurès; Boufarik; El Outaya; Hauts-plateaux
de Ghardaïa; Corso; Kabylie des Babor; Kabylie du Djurdjura, Aït oranais; Kabylie du Djurdjura, col de Tirourda, 1700 m; Monts de
Idjer; Oran; Sidi-Ferruch. Ros et al. (1990): Oued Rhiou. Boukha- Constantine; Monts de Tlemcen, Mazer; Monts de Tlemcen, Tiaret;
tem et al. (2017): El-Kala National Park, Tonga lake watershed. Ouarsenis, djebel Doui; Vallée du Chéliff, Duperré. Jelenc (1956,
Mazari & Boulaacheb (2023): Setif, Megriss Mountain. Frequent. 1967): Atlas Mitidjien, pic Mouzaïa. Bizot (1973): Djebel Belez-
ma à 15 km à l’ouest de Batna. Boukhatem et al. (2017): El-Kala
National Park, Tonga lake watershed. Frequent.
Streblotrichum enderesii (Garov.) Loeske
Barbula enderesii Garov. Syntrichia montana Nees var. calva
Nomenclature follows Kučera et al. (2013). (Durieu & Sagot ex Bruch & Schimp.) J.J.Amann

Records. — Jelenc (1955a): Philippeville; Stora. Rare. Records. — Jelenc (1955b, 1967): Mila. Mazari & Boulaacheb
(2023): Setif, Megriss Mountain. Rare.

Syntrichia caninervis Mitt. var. caninervis Syntrichia papillosissima (Copp.) Loeske

Records. — Trabut (1927), Jelenc (1954a, 1955a), Kramer (1980): Records. — Gallego (2005): Ancien départament de Constan-
Beni-Ounif. Very rare. tine. Boukhatem et al. (2017): El-Kala National Park, Tonga lake
watershed. Rare.

Syntrichia fragilis (Taylor) Ochyra

Syntrichia princeps (De Not.) Mitt.
Records. — Gauthier (1987): Assekrem, Hoggar. Very rare.
Records. — Bescherelle (1882): Petit Atlas. Bescherelle (1882),
Jelenc (1955a): Beni-Salah, sur les cèdres; Djebel Babor et Tababor;
Monts Batna, djebel Aurès. Brongniart (1882): Blidah. Corbière
Syntrichia handelii(Schiffn.) S.Agnew & Vondr. (1904): Médéa. Corbière (1904), Jelenc (1955a): Batna, flancs sud
et nord du djebel Tougourth. Jelenc (1953, 1955a, 1967): Massif
Records. — Pierrot (2000): Massif de l’Aizer, 1900 m. Boukhatem de l’Ouarsenis, cédraie de Téniet el Had, 1500 m. Jelenc (1953,
et al. (2017): El-Kala National Park, Tonga lake watershed. Rare. 1967): Massif de l’Aurès, flanc nord du djebel Pharaoun, 1800 m;

100 CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5)

 Revisiting the Bryophyte Checklist of Algeria

Massif de l’Aurès, forêt de Sgag, 1700 m. Jelenc (1955a): Atlas de **Tetrastichium fontanum (Mitt.) Cardot
Blida, gorges de la Chiffa; Kabylie du Djurdjura, Fort National.
Jelenc (1967): Massif du Bou Zegza, gorges de Keddara, 400 m.
Kramer (1980): Kabylie Mtns., Col de Tegma, Yakouren near
Comment
Azazga. Frequent. Excluded from Algeria. It was reported by Jelenc (1955a) from
Tassili n’Adrar, point d’eau Tig Hert, à l’ouest de Tamada,
600 m, but in the Addenda of the publication, the author
Syntrichia ruraliformis (Besch.) Mans rectified the identification to Vesicularia reimersiana Bizot &
P. de la Varde.
Syntrichia ruralis var. ruraliformis (Besch.) Delogne

Nomenclature follows Hedenäs et al. (2019). Thamnobryum alopecurum (Hedw.) Gangulee

Records. — Corbière (1904), Jelenc (1955a): Médéa; Sétif, 1120 m. Records. — Bescherelle (1882): Djurjura. Bescherelle (1882),
Jelenc (1955a): Hauts-plateaux oranais; Mila; Tala Ghizan. Kramer Corbière (1904), Jelenc (1955a): Gorges de la Chiffa, ruisseau des
(1980): Djebel Belezma, 15 km W de Batna. Mazari & Boulaacheb Singes. Bescherelle (1882), Jelenc (1955a): Bône; Édough; La Calle;
(2023): Setif, Megriss Mountain. Rare. Philippeville; Stora. Trabut (1888): sine loco. Dubuis & Faurel (1945):
Kabylie du Djurdjura, plateau de l’Aizer, 1900 m. Feldmann (1947),
Jelenc (1955a, 1959): Kabylie de Collo, Baie de Tamanart, moins
Syntrichia ruralis (Hedw.) F.Weber & D.Mohr de 50 m; Kabylie de Collo, vallée de l’oued Bin el Ouidane, près
de Taoulel, 600 m. Jelenc (1951, 1967): Monts de Tlemcen, forêt
de Zariflet, 1100-1200 m. Jelenc (1953, 1955a, 1967): Kabylie du
Records. — Bescherelle (1882): Djurdjura; Kabylie; Petit Atlas.
Djurdjura, massif de l’Aizer, flanc nord du djebel Tachgagalt, 2000 m.
Bescherelle (1882), Jelenc (1955a): Constantine, sommet du
Jelenc (1955a): Arba; Azerou d’Aït Zikki, Ifri bou Ouadufel; Kabylie
Mansourah; Massif de la Mahouna, Serdj el Aouda; Oran, hauts
de Collo; Kabylie du Djurdjura, Aït Ouaban; Kabylie du Djurdjura,
plateaux; Tiaret, 1000 m. Brongniart (1882), Jelenc (1955a),
Akfadou; Kabylie du Djurdjura, Azerou Goujane; Kabylie du Djurd-
Ahayoun et al. (2007): Atlas de Blida. Trabut (1887a): D’Oran à
jura, Tala Ghizan. Jelenc (1967): Monts de Tlemcen, forêt de Zariflet,
Mécheria. Corbière (1904): Tiaret, Hauts-Plateaux, Douar Tor-
1100-1200 m; Sahel de Koléa, ravin des Voleurs. Boukhatem et al.
rich, 1000 m. Corbière (1904), Jelenc (1955a): Médéa. Corbière
(2017): El-Kala National Park, Tonga lake watershed. Common.
(1905), Jelenc (1955a): Environs de Constantine, djebel Ouach.
Jelenc (1949, 1967): Haute plaine de Tlemcen, berges du chabet
bel Abbès, 750 m; Monts de Tlemcen, Forêt de Tessera Mramet,
1350-1400 m; Jelenc (1949, 1951, 1967): Monts de Tlemcen, **Thuidium assimile (Mitt.) A.Jaeger
escarpements du djebel el Béniane, 1100-1150 m. Jelenc (1951):
Djebel el Hassi, 1100 m. Jelenc (1951, 1967): Massif du Tessala, Comment
pic de Tafaraoui, 650 m. Jelenc (1953, 1955a, 1967): Kabylie du
Djurdjura, Aizer, flanc nord du djebel Tachgagalt, 1950 m. Jelenc
Excluded from Algeria. It was cited in error for Algeria by Ros
(1953, 1967): Bèlezma, crêtes de Bordjem, 2200 m; Kabylie Ba- et al. (1999) under the name T. philibertii Limpr.
bor, flanc nord du Babor, 1700 m; Massif de l’Aurès, flanc nord
du Chélia, 1800 m; Massif de l’Aurès, flanc nord-ouest du Ché-
lia, 1500 m; Massif de l’Aurès, flanc nord-ouest du Ras Keltoun, Thuidium recognitum (Hedw.) Lindb.
2000 m; Massif du Bélezma, cédraies de l’oued Chaba, 1400 m.
Jelenc (1955a): Akfadou; Aumale; Azéba; Col de Tirourda, 1700 m; Records. — Bescherelle (1882), Jelenc (1955a): Kabylie du Djurd-
Corso; Djebel Chenoua; Kabylie des Babor, cap Aokas; Kabylie jura, Tizi Ouzou. Very rare.
du Djurdjura, Fort National, 1000 m; Lambèse; Monts de Tlem-
cen, Mazer. Jelenc (1967): Haute-Plaine de Tlemcen, Chabet
bel-Abbès, 750 m. Boukhatem et al. (2017): El-Kala National Timmia bavarica Hessl.
Park, Tonga lake watershed. Mazari & Boulaacheb (2023): Setif,
Megriss Mountain. Common. Records. — Trabut (1914), Jelenc (1955a): Kabylie du Djurdjura,
forêt des Aït-Ali. Jelenc (1948, 1955a): Kabylie des Babor, Tababor.
Jelenc (1953): Flanc nord du djebel Tachgagalt, 2000 m. Jelenc
Syntrichia subpapillosissima (1953, 1955a): Kabylie du Djurdjura, Aizer, parois rocheuses des
(Bizot & R.B.Pierrot ex W.A.Kramer) puits de neige, 1900 m. Jelenc (1955a): Kabylie du Djurdjura, Aït
M.T.Gallego & J.Guerra Daoud; Kabylie du Djurdjura, Azerou Tidjer, 1300 m; Kabylie du
Djurdjura, crêtes du Djurdjura; Kabylie du Djurdjura, Ifri Maarès;
Kabylie du Djurdjura, Tizi Boussouil, 1800 m. Rare.
Records. — Bizot (1973): Fontaine Chaude à 30 km NE de Batna.
Bizot (1973), Kramer (1980): Djebel Belezma à 15 km à l’ouest
de Batna. Rare. Timmia megapolitana Hedw.
Records. — Bescherelle (1882): Djebel Goufi; Djurjura, Djebel
Syntrichia virescens (De Not.) Ochyra Tababor. Lapie (1909): Djurjura. Rare.

Records. — Jelenc (1955a): Kabylie du Djurdjura, Fort National,

1000 m. Boukhatem et al. (2017): El-Kala National Park, Ton- Comment
ga lake watershed. Mazari & Boulaacheb (2023): Setif, Megriss There is confusion about the presence of the species of Timmia
Mountain. Rare. in Algeria, and even in northern Africa due to the taxonomical

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 Ros R. M. et al.

difficulties in separating T. megapolitana and T. bavarica. Tlemcen, forêt de Tessera Mramet, 1375 m. Jelenc (1955a): Boghar;
Jelenc (1955a) considered both species synonyms, joining Ouarsenis. Rare.
all the previous reports of the genus under T. bavarica. As
Ros et al. (1999, 2013) did not realize of the old reports of
T. megapolitana, this species was ignored also in both check- Tortella inclinata (R.Hedw.) Limpr.
lists. After Brassard (1984) they are considered different taxa.
Unfortunately, no habitat information in Algerian literature Records. — Corbière (1904), Jelenc (1955a): La Calle. Corbière
has been given that could help to distinguish both species. (1905), Jelenc (1955a): Environs de Constantine, djebel Ouach.
Jelenc (1955a): Alger; Hammam Meskoutine. Boukhatem et al.
The presence and distribution of Timmia species in Algeria (2017): El-Kala National Park, Tonga lake watershed. Rare.
should be reevaluated.
Comment
Tortella inclinata var. brachypoda (Besch.) Paris was described
Timmiella barbuloides (Brid.) Mönk. by Bescherelle (1882) from Alger as Barbula inclinata var.
brachypoda Besch. and later compiled in Jelenc (1955a)
Records. — Montagne (1838), Bescherelle (1882), Corbière (1904): but according to Ros et al. (2013), its identity has not been
Alger. Bescherelle (1882): Géryville. Bescherelle (1882), Corbière revised since.
(1904), Jelenc (1955a): Bône, vieux mur au pied de l‘Édough. Be-
scherelle (1882), Jelenc (1955a): Constantine; Djebel Ksel; Oran;
Philippeville. Gandoger (1883): sine loco. Trabut (1887a): D‘Oran
à Mécheria. Corbière (1889): Blida, Route de Dalmatie. Cardot Tortella inflexa (Bruch) Broth.
(1904), Jelenc (1955a): Environs d‘Alger, Bouzarea. Corbière (1904),
Jelenc (1955a): Médéa; Orléansville. Corbière (1904), Jelenc (1955a), Records. — Bescherelle (1882): Mustapha-Supérieur. Bescherelle
Ros et al. (1990): Gorges de la Chiffa, ruisseau des Singes. Corbière (1882), Jelenc (1955a): Alger, rochers calcaires de Birmandreïs;
(1905): Environs de Constantine, ravin du Séminarie; Environs de Tixeraïn. Corbière (1904), Jelenc (1955a): Camp des Chênes, rive
Constantine, Sidi M’Cid. Camus (1906), Jelenc (1955a): environs droite de la Chiffa; Orléansville, bois de la Pépinière. Rare.
de Tlemcen. Lapie (1909): Kabylie du Djurjura. Jelenc (1951,
1967): Massif du Tessala, pic de Tafaraoui, 500 m; Sahel d’Arzeu,
djebel Borousse. Jelenc (1955a): Aït Khalfoun; Atlas de Blida, oued Tortella nitida (Lindb.) Broth.
el Kebir; Djebel Amour; Kabylie du Djurdjura, Fort National. Jelenc
(1955a, 1956, 1967): Sahel d’Alger, dans le ravin du Frais-Vallon.
Jelenc (1956, 1967): Atlas Mitidjien, pic Mouzaïa. Jelenc (1956, Records. — Corbière (1889, 1904), Jelenc (1955a): Blida, grèves
1967): Massif de la Bouzaréa, forêt de Baïnem. Jelenc (1967): Massif de l’oued Beni-Aza. Corbière (1904), Jelenc (1955a): Oran, Mont
du Bou Zegza, gorges de Keddara, 400 m; Zahrez Chergui, dejbel Santa-Cruz. Corbière (1904), Jelenc (1955a): Mostaganem. Corbière
Touilila, 1000 m. Ros et al. (1990): Between Mostaganem and Cap (1905), Jelenc (1955a): Environs de Constantine, Sidi M’Cid. Lapie
Ivi, Abdelmaimalek-Ramdan beach, 0 m; Djebel Chenoua, 5-40 m; (1909), Jelenc (1955a): Kabylie du Djurjura. Maire (1916), Jelenc
Hammam-Riga, 500 m; Texenna’s wood, 700 m. Ahayoun et al. (1955a): Sud oranais, Monts des Ksour, le djebel Aïssa, 2250 m.
(2007): Mustapha supèrieur. Boukhatem et al. (2017): El-Kala Jelenc (1949, 1967): Monts de Tlemcen, col du Juif, 800 m. Jelenc
National Park, Tonga lake watershed. Common. (1951, 1967): Massif du Tessala, pic de Tafaraoui, 650 m. Jelenc
(1955a): Alger, Atlas de Blida, oued el Kébir; Édough; Touggourt.
Boukhatem et al. (2017): El-Kala National Park, Tonga lake watershed.
Mazari & Boulaacheb (2023): Setif, Megriss Mountain. Frequent.
Tortella flavovirens (Bruch) Broth.
Comment
Records. — Bescherelle (1882), Jelenc (1955a): Alger; Bône, dunes Tortella nitida var. subtortuosa (Boulay) Jelenc was reported
des Beni-Urgin, sur les genévriers; Bouzaréah; Kabylie du Djurdjura; by Corbière (1904) from Oran, Mont Santa-Cruz and Cor-
La Calle; Philippeville. Corbière (1904), Jelenc (1955a): Monts de bière (1905) from the environs de Constantine, Sidi M’Cid
Batna, flanc sud du djebel Tougourth, 1700 m. Jelenc (1955a): Aïn
Taya; Alma; Atlas de Blida; Cap Matifou; Cherchell; Corso; Djebel as Barbula nitida var. subtortuosa (Boulay) Corb. and later
Murdjadjo; Kabylie du Djurdjura, djebel Gouraya; Marengo; Oued compiled in Jelenc (1955a) but according to Ros et al. (2013)
el Kébir, 200 m; Sidi-Ferruch. Jelenc (1956, 1967): Monts du Dahra, its identity has not been revised since. The search of the type
au Cap Ivi. Jelenc (1967): Sahel de Koléa, Castiglione. Ros et al. has not been fruitful (Ros, unpublished). Ros et al. (2013)
(1990): Between Mostaganem and Cap Ivi, Abdelmaimalek-Ramdan mentioned erroneoulsy that Tortella nitida var. media (Boulay)
beach, 0 m; Cape Falcon, les Andalouses, 30-50 m; Dellys, 0-40 m;
Skikda beach, 0 m. Boukhatem et al. (2017): El-Kala National Park, Corb. were reported from Algeria.
Tonga lake watershed. Mazari & Boulaacheb (2023): Setif, Megriss
Mountain. Common.
Tortella squarrosa (Brid.) Limpr.
Records. — Bescherelle (1882): Bône; Constantine; Kabylie; Petit
Tortella humilis (Hedw.) Jenn. Atlas. Bescherelle (1882), Jelenc (1955a): Alger, au dessus du Fort
l‘Empereur; Batna; Djebel Babor; Oran; Tiaret. Gandoger (1883):
Records. — Bescherelle (1882), Jelenc (1955a): Alger; Birkadem; sine loco. Trabut (1887a): D‘Oran à Mécheria. Brongniart (1882),
Constantine, fissures des rochers du djebel Chettaba. Gandoger Corbière (1889): Blida. Corbière (1904), Corbière (1905): Cons-
(1883): sine loco. Corbière (1905), Jelenc (1955a): Environs de tantine, bois du djebel Ouasch, 1100 m. Corbière (1904), Jelenc
Constantine, djebel el Hadji Baba. Jelenc (1949, 1967): Monts de (1955a): Médéa; Orléansville, bois de la Pépinière. Lapie (1909):

102 CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5)

 Revisiting the Bryophyte Checklist of Algeria

Kabylie du Djurjura. Thériot & Trabut (1930), Jelenc (1955a), Cano Tortula acaulon var. retortifolia (J.Guerra & Ros)
et al. (2000): Montium Aurasiorum prope Calceum Herculis, El R.H.Zander
Kantara. Jelenc (1949, 1967): Djebel Murdjadjo (Oran), ravin de
l’oued Tammermouth; after the name used by the first author has Records. — Guerra et al. (1991): Tichi. Very rare.
been put and in parentheses. Jelenc (1951, 1967): Massif du Tessala,
pic de Tafaraoui, 650 m. Jelenc (1954b): Atlas mitidjien: fôret d’el
Aioun, au Sud de Bouinan, 500 m. Jelenc (1954b, 1967): Flanc
nord du Sahel de Koléa (Alger), Castiglione, dune de la Station Tortula atrovirens (Sm.) Lindb.
expérimentale d’Aquiculture at de Pêche; Massif du Bou Zegza,
gorges de Keddara, 400 m. Monjauze (1954), Jelenc (1967): Forêt
de Guetarnia, vallon du Chabet el Hamra, 680 m. Jelenc (1955a): Records. — Bescherelle (1882): Djurjura. Bescherelle (1882),
Atlas de Blida, oued el Kebir, 200 m; Azéba; Djebel Tafrent; El Corbière (1889), Jelenc (1955a): Blidah. Bescherelle (1882),
Guerrah; Gorges de la Chiffa, 300-400 m; Kabylie du Djurdjura, Jelenc (1955a): Alger, pied des rochers, couteaux Bab-Azoun;
Fort National, 1000-1200 m; Monts de Constantine, djebel Oua- Bouzaréah; Oran. Trabut (1887a): D’Oran à Mécheria. Corbière
ch, Hammam Meskoutine; Monts de Tlemcen, Sidi Djillali. Jelenc (1904), Jelenc (1955a): Médéa; Mostaganem, pente sablonneuse
(1956, 1967): Massif du Tessala, pic Tessala, 1000 m. Bizot (1973): descendant à la Marine. Corbière (1905), Jelenc (1955a): En-
Fontaine Chaude à 30 km NE de Batna; Ruines de Timgad. Ros virons de Constantine, djebel Ouach. Camus (1906), Coppey
et al. (1990): Oued Rhiou; Ouled Mimoum. Mazari & Boulaacheb (1911), Jelenc (1955a): Ben Zireg, dans l’extrème-Sud Oranais.
(2023): Setif, Megriss Mountain. Common. Thériot (1931, 1933): Hoggar, in montibus Atakor-n-Ahaggar,
2000 m. Thériot (1931, 1933), Jelenc (1955a): Hoggar, djebel
Amezzeroui, 2600 m. Jelenc (1955a): Biban; Hoggar, llaman,
2000 m; Kabylie du Djurdjura, Fort National, 1000-1200 m;
Tortella tortuosa (Hedw.) Limpr. Monts de Constantine; Ouarsenis. Collenot et al. (1960): Tassi-
li N’Aijer, Afaïtalèsse. Jelenc (1967): Atlas Mitidjien, forêt d’El
Aioun, 650 m; Hoggar. Ros et al. (1990): Dellys, 0-40 m; Ouled
Tortella tortuosa var. fragilifolia (Jur.) Limpr. Mimoum. Common.
Nomenclature follows Hodgetts et al. (2020).
Records. — Bescherelle (1882): Sud d’Oran. Trabut (1887a): Tortula canescens Mont.
D’Oran à Mécheria, Tell. Jelenc (1951): Koudiat d’Hafir, 1250 m;
Monts de Tlemcen: forêt de Tessera Mramet, 1400 m. Jelenc (1951, Records. — Corbière (1905), Jelenc (1955a): Environs de Cons-
1967): Monts de Tlemcen, djebel el Béniane, 1050 m.Jelenc (1953, tantine, djebel Ouach. Jelenc (1955a): Atlas de Blida, oued el Kebir,
1967): Aurès, djebel Lazreg, 1900 m; Aurès, flanc nord du Chélia, 200 m; Aumale; Tiaret. Boukhatem et al. (2017): El-Kala National
1700 m; Aurès, flanc sud du Chélia, 1700 m; Aurès, Ras-el-Ma, forêt Park, Tonga lake watershed. Rare.
de Sgag, 1700 m. Jelenc (1955a): Aurès; Constantine; Djurdjura;
Hauts-plateaux oranais; Kabylie du Djurdjura, col de Tirourda,
1700 m; Littoral constantinois; Monts de Tlemcen, Mazer; Monts
de Tlemcen, Tlemcen. Jelenc (1956, 1967): Massif du Tessala, pic Tortula caucasica Broth.
Tessala, 1000 m. Jelenc (1966, 1967): Zahrez Chergui, oued M’Kam-
za. Jelenc (1967): Monts de Tlemcen, 1200-1400 m. Boukhatem Records. — Bescherelle (1882), Jelenc (1955a): Constantnie, sur
et al. (2017): El-Kala National Park, Tonga lake watershed. Frequent. les rochers couverts de terre, chute du Rummel; Oran, près Mer-
s-el-Kebir; Philippeville. Braun-Blanquet & Maire (1924), Jelenc
(1955a): Tlemcen. Jelenc (1955a): Gorges de la Chiffa; Oran. Rare.
Comment
Köckinger & Hedenäs (2023) identified eight morphologically
and molecularly distinct entities at the species level within Tortula cuneifolia (Dicks.) Turner
the former T. tortuosa. The identity and distribution of the
Records. — Montagne (1838), Bescherelle (1882), Jelenc (1955a):
T. tortuosa complex in Algeria should be reevaluated in accord- Alger. Bescherelle (1882): Philippeville. Bescherelle (1882), Jel-
ance with the new taxonomic criteria. enc (1955a): Bouzaréah; El Arrouch. Bescherelle (1882), Jelenc
(1955a), Cano et al. (2000): La Calle, parois verticales des fos-
sès, forêt du lac Touga (Tonga); Bône. Corbière (1904), Jelenc
(1955a): Kabylie du Djurdjura, Fort National, 1000 m. Corbière
Tortula acaulon (With.) R.H.Zander (1905), Jelenc (1955a): Environs de Constantine, djebel Ouach.
Lapie (1909): Kabylie du Djurjura. Jelenc (1955a): Aïn Taya;
Records. — Bescherelle (1882), Jelenc (1955a): Alger; Constantine, Atlas de Blida, oued el Kebir, 200 m; Kabylie du Djurdjura, col
collines de la Casbah; Oran. Corbière (1904), Jelenc (1955a): Sétif, des Aït Aïcha; Kabylie du Djurdjura, Isser; Kabylie du Djurd-
1112 m. Jelenc (1951, 1967): Haute-Plaine de Tlemcen, prés le long jura, pic Belloua; La Réghaia; Monts de Tlemcen, Tlemcen, forêt
de l’oued el Horra, au pied des collines de Bréa. Rare. d’Hafir; Ouarsenis, Téniet el Had. Jelenc (1956, 1967): Atlas
Mitidjien, pic Mouzaïa. Jelenc (1967): Kabylie du Djurdjura,
Yakouren, 1000 m; Massif du Bou Zegza, gorges de Keddara,
Tortula acaulon var. pilifera (Hedw.) R.H.Zander 400 m. Frequent.

Records. — Bescherelle (1882), Jelenc (1955a): Alger; Constantine,

collines de la Casbah; Oran. Trabut (1887a): D’Oran à Mécheria. Tortula freibergii Dixon & Loeske
Corbière (1905): Environs de Constantine, route de Batna; Envi-
rons de Constantine, Salah Bey. Braun-Blanquet & Maire (1924), Records. — Pierrot (2000): Environs d’Alger, dans le Frais Vallon,
Jelenc (1955a): Tlemcen. Rare. près d’El Biar. Very rare.

CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5) 103

 Ros R. M. et al.

Tortula inermis (Brid.) Mont. du Djurdjura, Tizi Ouzou. Gandoger (1883), Ahayoun et al. (2007):
sine loco. Trabut (1887a): D’Oran à Mécheria. Corbière (1889),
Records. — Bescherelle (1882): Petit Atlas; Sahara algérien. Jelenc (1955a): Blida. Corbière (1904), Jelenc (1955a): Cherchell,
Bescherelle (1882), Corbière (1905): Constantine; Environs de sur des ruines, chemin du cap Zizerine; Kabylie du Djurdjura, Fort
Constantine, Chettabah. Bescherelle (1882), Jelenc (1955a): Alger; National, 1000 m; Sétif, 1080 m. Corbière (1905): Environs de
Bou-Sâada; Monts de Tlemcen, Tlemcen; Monts des Trara, Col Constantine, Le Hamma; Environs de Constantine, Les Pins; En-
de Nédroma; Oran, hauts plateaux; Sud algerois, Gueelt es Stel. virons de Constantine, Menidj; Environs de Constantine, ravin du
Trabut (1887a): D’Oran à Mécheria. Corbière (1904): Tlemcen, Rummel; Environs de Constantine, route de Sétif. Corbière (1905),
sentier de Bou-Médine à la cascade. Corbière (1905): Environs de Jelenc (1955a): Environs de Constantine, Sidi M’Cid. Corbière
Constantine, Les Pins; Environs de Constantine, Menidj; Environs (1905), Jelenc (1966, 1967): Environs de Constantine, route du
de Constantine, ravin du Séminaire; Environs de Constantine, route djebel Ouach. Camus (1906), Braun-Blanquet & Maire (1924):
de la Corniche; Environs de Constantine, route de Sétif; Environs de Environs de Tlemcen. Maire & Sevenet (1928), Jelenc (1955a):
Constantine, Sidi M’Cid. Camus (1906): Oran. Braun-Blanquet & Tombeau de la Chrétienne, près de Bérard. Jelenc (1949, 1967):
Maire (1924): Tlemcen, 800 m. Szepesfalvi (1932), Andreánszky Monts et Haute-Plaine de Tlemcen. Jelenc (1951, 1967): Massif
(1934), Jelenc (1955a): Aflou, in monte djebel Sidi Okba, 1700 m. du Tessala, pic de Tafaraoui, 650 m. Jelenc (1953, 1967): Aurès,
Jelenc (1951, 1967): Monts de Tlemcen, djebel el Hassi, 1100 m. flanc nord du Ras Pharaoun, 1800 m; Bèlezma, cédraies de l’oued
Jelenc (1953): Massif de Guethiane, Kef Tachrirt, 1850 m. Jelenc Chaba, 1400 m. Jelenc (1955a): Atlas de Blida, oued el Kebir, 200 m;
(1953, 1967): Massif de l’Ouarsenis, pente du pic Sidi Abdelkader, Batna; Cap Aokas; Djebel Tessala; Djemila; Kabylie des Babor; La
1500 m. Jelenc (1955a): El Kantara, 1500 m; Kabylie des Babor, Bouzaréa; Marnia; Mascara; Médéa; Monts de Constantine; Oran;
cap Aokas; Kabylie des Babor, Dra el Arba; Kabylie du Djurdjura, Ouarsenis, djebel Doui; Rouiba. Collenot et al. (1960): Tassili N’Aijer,
Fort National, 1000 m; Mila; Monts de Constantine; Monts de Afaïtalèsse. Ahayoun et al. (2007): Oran, aux “Planteurs”. Maza-
Tlemcen, Mazer; Oran; Ouarsenis, Téniet el Had. Jelenc (1955a), ri & Boulaacheb (2023): Setif, Megriss Mountain. Very common.
Ahayoun et al. (2007): Atlas de Blida, oued el Kebir, 200 m. Jelenc
(1967): Gorges de Keddara, 400 m; Massif de Guethiane, Kef
Tachrirt, 1500 m; Massif du Bou Zezga; Massif du Zaccar; Zahrez Tortula muralis subsp. muralis var. aestiva
Chergui, djebel Touilila, 1100 m; Zahrez Chergui, oued M’Kamza, Brid. ex Hedw.
Ain Hallouf. Bizot (1973): Djebel Belezma à 15 km à l’ouest de
Batna. Mazari & Boulaacheb (2023): Setif, Megriss Mountain. Records. — Corbière (1905): Environs de Constantine, près du
Very common. Rummel. Jelenc (1955a): Alger; Djebel Dira; Kabylie du Djurdjura,
Fort National, 1000 m; Monts de Constantine. Jelenc (1956, 1967):
Massif de la Bouzaréa, dans la forêt de Baïnem. Rare.
Tortula lindbergii Broth.

Records. — Bescherelle (1882): Djurjura. Bescherelle (1882), Tortula muralis subsp. obtusifolia (Schwägr.) Culm.
Jelenc (1955a): Constantine. Trabut (1887a): D’Oran à Mécheria.
Corbière (1904), Jelenc (1955a): Sétif, colline pierreuse à l’ouest de Records. — Bescherelle (1882), Jelenc (1955a): Tiaret. Jelenc
la ville, 1000-1100 m. Corbière (1905): Environs de Constantine, (1955a): Boghni; Djurdjura. Jelenc (1955b): sine loco. Rare.
Menidj; Environs de Constantine, ravin du Séminarie. Corbière
(1905), Jelenc (1955a): Environs de Constantine, ravin du Rum-
mel. Jelenc (1955a): Kabylie du Djurdjura, Aït Idjer; Kabylie du Tortula revolvens (Schimp.) G.Roth
Djurdjura, Fort National; Littoral oranais. Jelenc (1955b, 1967):
Col de Sfa, près de Biskra. Jelenc (1966, 1967): Zahrez Chergui,
oued Mlilia au djebel Toulila, 1000 m. Frequent. Records. — Corbière (1905), Jelenc (1955a): Environs de Con-
stantine, djebel Chettabah. Jelenc (1951): Tell oranais, forêt de Mou-
lay Ismael, 110-120 m. Jelenc (1955a): Alger; Ben Chicao; Blida;
Médéa. Collenot et al. (1960): Tassili N’Aijer, Afaïtalèsse. Ros &
Tortula marginata (Bruch & Schimp.) Spruce Guerra (1987a): El Kantara. Ros et al. (1990): Oued Rhiou. Rare.

Records. — Bescherelle (1882): Birtraria. Bescherelle (1882), Jel-

enc (1955a): Alger, vieux murs; Blidah; Bône, parois des citernes;
Constantine, chute du Rummel; Mostaganem; Philippeville. Trabut Tortula schimperi M.J.Cano, O.Werner & J.Guerra
(1887a): D’Oran à Mécheria. Corbière (1904, 1905): Constantine,
vieux mur entourant la piscine de Sidi M’Cid. Corbière (1905): En- Records. — Bizot (1973): Djebel Belezma à 15 km à l’ouest de
virons de Constantine, Moulin Laire. Jelenc (1951, 1967): Monts Batna. Very rare.
de Tlemcen, berges de l’oued Khémis au Khémis, 800 m. Jelenc
(1955a): Frais Vallon; Kabylie du Djurdjura, Aberkan; Kabylie du
Djurdjura, Agoulmin; Kabylie du Djurdjura, Aït Idjer; Oran. Jelenc Tortula solmsii (Schimp.) Limpr.
(1966, 1967): Monts de Saida, cascades de Tifrit. Ahayoun et al.
(2007): Hydra. Boukhatem et al. (2017): El-Kala National Park, Records. — Jelenc (1955a): Alger. Very rare.
Tonga lake watershed. Frequent.

Tortula subulata Hedw.

Tortula muralis Hedw.
Records. — Bescherelle (1882): Sommet de l’Atlas; Sud d’Oran.
Records. — Montagne (1838), Bescherelle (1882), Jelenc (1955a): Bescherelle (1882), Lapie (1909): Djurjura. Bescherelle (1882), Jelenc
Alger. Bescherelle (1882): Mustapha; Philippeville. Bescherelle (1882), (1955a): Alger; Bordj-Boghni; Djebel Édough. Trabut (1887a), Jelenc
Jelenc (1955a): Birmandreïs, chemins creux; Constantine; Kabylie (1955a): D’Oran à Mécheria, Tell. Corbière (1889), Jelenc (1955a):

104 CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5)

 Revisiting the Bryophyte Checklist of Algeria

Atlas de Blida, près de la glacière Laval, 1200 m. Corbière (1904), Tortula wilsonii (Hook.) R.H.Zander
Jelenc (1955a): Kabylie, Fort National, Michelet, 1000-1200 m.
Corbière (1905), Jelenc (1955a): Environs de Constantine, djebel Records. — Bescherelle (1882): Alger; Blidah; Collines du Bou-
Ouach. Jelenc (1949, 1967): Monts de Tlemcen, Forêt de Tessera Ahmera et des Santons. Bescherelle (1882), Jelenc (1955a): Ravins
Mramet, 1350 m; Monts de Tlemcen, Forêt de Zariffet, 1100 m. du Bouzaréah. Jelenc (1955a): Atlas de Blida, oued el Kebir; Kabylie
Jelenc (1953): Aurès, flanc nord-ouest du Ras Keltoun, 2200 m; du Djurdjura, Bône; Kabylie du Djurdjura, Michelet. Jelenc (1955b,
Babor, flanc nord du Babor, 1700 m. Jelenc (1953, 1967): Atlas de 1967): Monts de Constantine, djebel Ouach. Rare.
Blida, Cèdraies du ravin Bleu à Chrèa; Aurès, flanc sud du Chélia,
1700 m. Jelenc (1955a): Hauts-plateaux oranais; Kabylie du Djurd-
jura, col de Tirourda, 1700 m; Kabylie du Djurdjura, Dra Inguel;
Monts de Ksour, djebel Aïssa; Monts de Tlemcen, Terni; Tiaret, Trichostomum brachydontium Bruch
1000-1100 m. Jelenc (1956, 1967): Massif du Zaccar. Jelenc (1967):
Aurès, Ras Kaltoun, 2200 m; Aurès, Ras-el-Ma; Massif du Babor, Records. — Bescherelle (1882), Jelenc (1955a): Gorges de la
Le Babor, 1300 m; Massif du Bou Zegza, gorges de Keddara, 400 m Chiffa, ruisseau des Singes. Jelenc (1955a): Aït Khalfoun; Alger;
Jelenc (1967), Ahayoun et al. (2007): Aumale. Mazari & Boulaacheb Atlas de Blida, oued el Kebir, 200 m; Corso; Kabylie du Djurdju-
(2023): Setif, Megriss Mountain. Common. ra, djebel Gouraya; Kabylie du Djurdjura, Fort National, 1000 m.
Jelenc (1967): Sahel d‘Alger, Haouch Khodja Berry, 150 m. Ros
et al. (1990): Azazga; Between Mostaganem and Cap Ivi, Abdel-
Tortula truncata (Hedw.) Mitt. maimalek-Ramdan beach, 0 m; Hammam-Riga, 500 m; Texenna’s
wood, 700 m. Boukhatem et al. (2017): El-Kala National Park,
Records. — Bescherelle (1882), Jelenc (1955a): Alma, champs Tonga lake watershed. Frequent.
sablonneux; Bône; La Calle. Trabut (1887a): D’Oran à Mécheria.
Corbière (1889): Blida. Corbière (1904), Jelenc (1955a), Ahayoun
et al. (2007): Alger. Jelenc (1955a): Baba Ali; Kabylie du Djurdjura, Comment
Fort National; Littoral oranais; Rouiba. Jelenc (1955a), Ahayoun Only the reports of T. brachydontium and T. mutabile Bruch
et al. (2007): Maison-Carrée. Frequent. ex De Not. nom. illeg. have been considered here for this spe-
cies. Ros et al. (2022) recognized besides T. brachydontium s.s.,
Comment another three species that had been considered as infraspecific
Pottia truncata var. archidioides (Besch.) Paris was described taxa (T. herzogii Ros, O.Werner & R.D.Porley and T. meridi-
by Bescherelle (1882) as P. truncata f. archidioides Besch. onale Ros, O.Werner & R.D.Porley) or synonymized with
from Oran, bord de la mare du djebel Santo. Jelenc (1955a) T. brachydontium (T. littorale Mitt.). The identity and distri-
compiled this report and added two localities (Maison-Carrée bution of the T. brachydontium complex in Algeria should be
and Rouiba). According to Ros et al. (2013), its identity has reevaluated following the new taxonomical criteria.
not been revised since.

Trichostomum littorale Mitt.

Tortula vahliana (Schultz) Mont.
Nomenclature follows Ros et al. (2022).
Records. — Bescherelle (1882), Jelenc (1955a): Alger; Tiaret,
1150 m. Corbière (1904): Alger, faubourgh de Telemly, talus. Records. — Feldmann (1947), Jelenc (1955a): Kabylie de Collo,
Corbière (1905), Jelenc (1955a): Environs de Constantine, Le vallée de l’oued Djerrah près du Melhab, 300-400 m. Jelenc (1955a):
Hamma; Environs de Constantine, près la tuilerie Moutin. Braun- Atlas de Blida, oued el Kebir, 200 m. Rare.
Blanquet & Maire (1924), Jelenc (1955a): Tlemcen. Bizot (1973):
Ruines de Timgad. Rare. Comment
See the previous comment. It was considered by Ros et al.
(1999, 2013) as a synonym of Trichostomum brachydontium.
Tortula viridifolia (Mitt.) Blockeel & A.J.E.Sm.
Records. — Corbière (1905), Jelenc (1955a): Environs de Con-
stantine, djebel Ouach. Very rare. Vesicularia reimersiana Bizot & P. de la Varde
Records. — Bizot & Potier de la Varde (1952), Jelenc (1955a):
Tortula wilczekii Meyl. Tassili n’Adrar, point d’eau Tig Hert, à l’ouest de Tamada, 600 m.
Very rare.
Records. — Meylan (1936), Jelenc (1955a): Insula Habiba majore.
Warnstorfia fluitans (Hedw.) Loeske
Comment
Meylan (1936) proposed the name Tortula muralis subsp. Records. — Bescherelle (1882): sine loco. Jelenc (1955a): Beni
mairei Meyl. but was interpreted as species level as Tortula Foughal; Maison-Carrée. Rare.
mairei Meyl. nom. illeg. later homonym by Jelenc (1955a), who
replaced it by Tortula wilczekii Meyl. However, the identity Weissia armata (Thér. & Trab.) Fedosov
of the taxon has not been revised because the type has not
been traced (Ros et al. 2013). Weissia condensa var. armata (Thér. & Trab.) M.J.Cano, Ros & J.Guerra

CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5) 105

 Ros R. M. et al.

Weissia papillosissima Laz. Weissia leptocarpa Schimp. ex Besch. hom. illeg.

Nomenclature follows Fedosov (2011). Records. — Paris (1894-1898): Algier? Jelenc (1955a): sine loco.
Records. — Thériot (1931, 1933), Jelenc (1955a), Cano et al.
(2000): Hoggar, oued Haman, 2000 m. Jelenc (1955a): Est de Ta- Comment
zerouk. Collenot et al. (1960): Tassili N’Aijer, Afaïtalèsse. Frahm The report of Jelenc (1955a) was explicitly based on Paris’
(1978): Laghouat. Rare.
Index Bryologicus distribution data (Paris 1894-1898: 1364).
Nevertheless, Paris gives as doubtful the presence in Africa,
Weissia brachycarpa (Nees & Hornsch.) Jur. as he gives “Alger?”, in addition to “Lusitania (Cintra)” in
Europe. As no original report has been found in the Alge-
Records. — Bescherelle (1882): Bougie; Édough; Stora. Besche- rian bryophyte flora literature, the report in the country and
relle (1882), Jelenc (1955a): Alger. Corbière (1905), Jelenc (1955a): northern Africa is doubtful. Furthermore it is an illegitimate
Environs de Constantine, route de la Corniche et près du Hamma. and doubtful name (Sérgio & Carvalho 2003).
Camus (1906), Jelenc (1955a): Oran; Tlemcen. Jelenc (1955b,
1967): Monts de Tlemcen, Mazer, dans les gorges de l’oued el
Khémis. Boukhatem et al. (2017): El-Kala National Park, Tonga
lake watershed. Rare. Weissia levieri (Limpr.) Kindb.

Records. — Brotherus (1924): Algier. Jelenc (1955a): sine loco.

Very rare.
Weissia condensa (Voit) Lindb.

Records. — Bescherelle (1882), Jelenc (1955a): Alger; Batna; Comment

Constantine, rochers du Rummel, Bougie. Corbière (1905), Jelenc Brotherus (1924) listed Algier in the distribution data of the spe-
(1955a): Environs de Constantine, Sidi M’Cid. Camus (1906), cies, and Jelenc (1955a) explicity based his report on Brotherus
Jelenc (1955a): Monts de Tlemcen; Oran. Jelenc (1955a): Djebel work, indicating “sine loco” instead of the city of Algier. Ros et al.
Chenoua; El Kantara; Kabylie du Djurdjura, Aizer; Kabylie du
Djurdjura, djebel Gouraya; Kef Mouella. Boukhatem et al. (2017): (1999, 2013), who based their lists on Jelenc (1955a) compila-
El-Kala National Park, Tonga lake watershed. Frequent. tion interpreted it as an imprecise locality in northern Africa and
was therefore not included in the Algerian list. Although the
report is imprecise and no original report has been found in the
Algerian bryophyte flora literature, the species could be present
Weissia controversa Hedw. in the country, as it is not a rare species in the Mediterranean.
R ecords . — Montagne (1838), Bescherelle (1882), Jelenc
(1955a): Alger. Bescherelle (1882): Blidah; Bône. Bescherelle
(1882), Jelenc (1955a): La Calle; Philippeville. Corbière (1889): Weissia longifolia Mitt.
Blida, bords d’un ruisselet, aux Chênes-Lièges. Corbière (1904),
Jelenc (1955a): Djebel Chenoua, La Bouzaréa. Corbière (1904, Records. — Bescherelle (1882): Alger, Mustapha, coteaux de l‘Ag-
1905), Jelenc (1955a): Constantine, bois du djebel Ouasch, ha. Bescherelle (1882), Jelenc (1955a): Bône; Philippeville. Jelenc
1100 m. Camus (1906), Jelenc (1955a): environs de Tlemcen. (1955a): Alger; Monts du Daya. Rare.
Lapie (1909): Kabylie du Djurjura. Jelenc (1955a): Atlas de Bli-
da, oued el Kebir, 200 m; Djebel Chenoua; Djidjelli; Kabylie
du Djurdjura, Afensou; Kabylie du Djurdjura, col de Tirourda, Comment
1700 m; Kabylie du Djurdjura, Fort National, 1000-1200 m. Astomum crispum var. minus Jelenc nom. inval. is based on the
Jelenc (1956, 1967): Massif du Bou Zegza, dans les gorges de
Keddara. Ros et al. (1990): Skikda beach, 0 m; Texenna’s wood, name Systegium crispum var. minus Besch. nom. nud. proposed
700 m. Frequent. by Bescherelle (1882) from Alger and Tiaret, and later compiled
by Jelenc (1955a). According to Ros et al. (2013), the original
material could not be found and therefore the identity of this
Weissia controversa var. crispata (Nees & Hornsch.) name has not been studied; if it merits recognition, then a
Nyholm new combination under Weissia longifolia may be required.
Records. — Jelenc (1955a): Kabylie du Djurdjura, col de Tirourda,
1700 m; Kabylie du Djurdjura, Fort National, 1000 m; Monts de
Constantine, djebel Ouach. Jelenc (1956, 1967): Atlas Mitidjien, Weissia multicapsularis (Sm.) Mitt.
pic Mouzaïa. Rare.
Records. — Corbière (1905), Jelenc (1955a): Environs de Con-
stantine, plateau du Mansourah. Very rare.
Weissia controversa var. densifolia (Bruch & Schimp.)
Wilson Weissia rutilans (Hedw.) Lindb.

Records. — Jelenc (1955a): La Bouzaréa; Monts de Constantine, Records. — Bescherelle (1882): Kabylie. Bescherelle (1882), Jelenc
djebel Ouach. Rare. (1955a): Édough, bord des chemins maures; Kabylie du Djurdjura,

106 CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5)

 Revisiting the Bryophyte Checklist of Algeria

Bouiroun; Philippeville. Gandoger (1883): sine loco. Jelenc (1955a): thallose liverworts, especially the Marchantiopsida, are the
Alger. Boukhatem et al. (2017): El-Kala National Park, Tonga lake best adapted (Bischler 2004).
watershed. Rare.
Among mosses, there is a clear dominance of acrocarpous
taxa (76.9% of the total number of mosses) over pleurocar-
pous taxa (23.1%). These data are also in agreement with
Weissia wimmeriana (Sendtn.) Bruch & Schimp. the data of other Mediterranean countries (Ros et al. 2013).
Regarding the dominant families (Table 2), within hornworts
Records. — Bescherelle (1882), Jelenc (1955a): Alger; Philippe- and liverworts, Ricciaceae is the best represented, with 31 taxa
ville. Rare.
(25.6% of the liverworts and hornworts’ taxa). The aquatic
and ephemeral Riellaceae, with eight taxa, is the second-best
Zygodon rupestris Schimp. ex Lorentz represented (6.6%), followed by Fossombroniaceae (seven taxa,
5.8%), Lophocoleaceae and Scapaniaceae (six taxa, 5.0%
Records. — Boukhatem et al. (2017): El-Kala National Park, each), and Cephaloziellaceae and Porellaceae (five taxa, 4.1%
Tonga lake watershed. Very rare.
each). The remaining 43.8% of hornwort and liverwort taxa
belong to 25 different families. Concerning mosses, Pottiaceae
Zygodon viridissimus (Dicks.) Brid. is the dominant family, with 107 taxa (28.4% of the moss
taxa), as it is well known for being the largest in the mosses
Records. — Bescherelle (1882): Blidah; Kabylie; Oran. Bescherelle in number of genera and consists of plants of small size well
(1882), Jelenc (1955a): Alger; Birmandreïs; Bouzaréah; Gorges de
la Chiffa; Hauts-plateaux oranais, Chellala Dahrania; La Calle, sur adapted to harsh environments (Zander 1993). It is followed
les chênes; Philippeville. Trabut (1887a): D’Oran à Mécheria. Jelenc at a great distance by Brachytheciaceae (39 taxa, 10.3%).
(1955a): Alma; Kabylie des Babor, cap Aokas; Khodjaberry; Littoral Much fewer taxa belong to Bryaceae (27 taxa, 7.2%), Grim-
oranais; Marengo. Frequent. miaceae (26 taxa, 6.9%), Orthotrichaceae (24 taxa, 6.4%),
Fissidentaceae (20 taxa, 5.3%), Mniaceae (16 taxa, 4.2%),
Comment Funariaceae (15 taxa, 4.0%), and Bartramiaceae (10 taxa,
Zygodon viridissimus var. vulgaris Malta fo. mediterranea Malta 2.7%). The remaining 24.7% of the moss taxa belong to
was cited by Jelenc (1949, 1967) from Sahel d’Alger, Ben 37 different families. Further analysis of the catalogue (Table 2)
Aknoun, 250 m. The contradictory combination assignation reveals several genera that are especially well represented. It is
of the two infraspecific taxa makes it not possible to assign not surprising that the genus Riccia is the most species-rich
any species to this name. As Ros et al. (1999, 2013) stated, in Algeria, with 30 taxa (24.8% of the liverwort and horn-
Zygodon viridissimus var. vulgaris Malta nom. illeg. is an old wort taxa), followed at a great distance by Riella (eight taxa,
name for Zygodon rupestris Schimp. ex Lorentz, and Zygodon 6.6%), Fossombronia (seven taxa, 5.8%), and Cephaloziella
viridissimus fo. mediterraneus Malta is a taxon related to Zygo- and Porella (five taxa, 4.1% each). Remarkably, most of them
don viridissimus (Dicks.) Brid. are annual or short-lived, and can develop during the short,
humid periods, both in the Mediterranean and Saharan areas.
The remarkable diversity of Riccia in the Mediterranean
DISCUSSION (Jovet-Ast 1986; Bischler 2004) is a well-known fact. But in
the Maghreb countries in general, and Algeria in particular,
Species diversity the genus Riella is especially diverse (Jelenc 1957; Ros et al.
Based on this catalogue, a total of 477 species, as well as 2013; Segarra-Moragues et al. 2014), probably due to the
21 infraspecific taxa, are reported from Algeria, resulting in abundance of steppe areas in the Hauts Plateaux algériens,
a total of 498 taxa (Table 1). They include three hornwort with many temporary lagoons.
species, 113 liverwort species (118 taxa), and 361 moss spe- Regarding mosses, the most species-rich genus is Tortula,
cies (377 taxa). with 22 taxa (5.8% of the moss taxa), followed by Fissidens
Compared to the previous Algeria checklists (Ros et al. (20 taxa, 5.3%); Ptychostomum (17 taxa, 4.5%), Grimmia
2007a, 2013), which included five taxa of hornworts, 121 of (16 taxa, 4.2%), Orthotrichum (13 taxa, 3.4%), Syntrichia
liverworts, and 352 of mosses, the current update indicates a (12 taxa, 3.2%), Weissia (11 taxa, 2.9%), Didymodon, and
decrease of two species of hornworts and three of liverworts, Entosthodon (ten taxa, 2.7% each).
alongside a net increase of 25 moss taxa.
The analysis of the floristic catalogue reveals that the Species richness
hornwort and liverwort taxa are scarce, representing 24.3% Analysis of Algerian species richness with other Maghreb coun-
of the total, whereas mosses represent 75.7%. Hornworts tries (Table 3) places Algeria as the country with the second
represent 0.6% of the bryophytes (2.5% of the total num- largest number of taxa. Having 498 taxa, Algeria is situated
ber of liverworts and hornworts together). Among hepat- behind Morocco, which contains 526, and Tunisia, which has
ics, foliose species represent 39.7%, and thallose is 57.8%. 305 taxa, is in the last place. Further analysis indicates that
These results are not surprising, as the hornworts grow in the percentages of the groups of bryophytes are similar. The
wet habitats (Frahm 2010) and are less-well adapted to the percentages of liverworts and hornworts together are between
climatic and edaphic Mediterranean conditions, whereas the 21.1 and 29.8% in the three countries (24.3% in Algeria,

CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5) 107

 Ros R. M. et al.

Table 1. — Species diversity data of bryophytes in Algeria.

Number of
Number of infraspecific Percentages in
species taxa Number of taxa (%) hornworts + liverworts group Percentages in mosses
Hornworts 3 0 3 (0.6%) 2.5% –
Liverworts 113 5 118 (23.7%) 97.5% including 39.7% foliose and –
57.8% thallose
Mosses 361 16 377 (75.7%) – 76.9% acrocarpous
23.1% pleurocarpous
Total 477 21 498 – –

21.1% in Morocco, and 29.8% in Tunisia). Moss percentages higher (6.72 × 10-3) than in any of the Maghreb countries,
are in the range of 70.2 and 78.9% (75.7% in Algeria, 78.9% which allows us to confirm that the degree of knowledge of
in Morocco, and 70.2% in Tunisia). Tunisia seems to have a the bryophyte flora in northern African countries, which pre-
higher percentage of liverworts and hornworts than Algeria sent physical environments, forests, and climatic conditions
and Morocco, despite having similar climatic conditions similar to those in the south of the European continent, is
in the coastal areas. This is probably due to the deep study still incomplete.
carried out by Jovet-Ast & Bischler (1971), who visited the
country and made an exhaustive study of these plants. Even Species abundance
so, it was amplified recently by Ben Osman et al. (2019) by According to the relative abundance of species in Algeria,
studying the Kroumiria region, in northern Tunisia. About 98 taxa (19.7%) are very rare, 263 taxa (52.8%) are rare,
mosses, the higher number of mosses in Morocco is clearly 77 (15.5%) are frequent, 48 (9.6%) are common, and only
due to having been the object of much more attention and 12 (2.4%) are very common. These data indicate that 72.5%
publications in the last decade of the 20th century and the of the catalogued taxa are rare and very rare, and 27.5% are
first decade of the 21st century by Spanish bryologists of frequent, common, and very common. All the abundance
the University of Murcia and the Autonomous University categories surprisingly are present in very similar percent-
of Madrid. Even lately, Moroccan researchers from the Uni- ages in Algeria as in studies in other areas, e.g. in the Sierra
versity of Ibn Tofail (Kenitra) are making many interesting Nevada Mountains, in southern Spain (Rams et al. 2014).
additions to the country (e.g. Zaza et al. 2018; Laouzazni That can lead to the assumption that the deviation due to the
et al. 2020; Saadi et al. 2020). In Europe, the bryophytic flora undercollection in Algeria is similar to other Mediterranean
is much better known than in the Maghreb. Countries like areas. Only 12 mosses are very common, but surprisingly some
Greece, Italy, and Spain, neighboring countries of Algeria, very common species in other Mediterranean areas (Ros &
have a greater bryophytic richness, containing 743, 1240, Guerra 1987b; García-Zamora et al. 1998; Rams et al. 2014;
and 1182 taxa, respectively. The liverwort and hornwort Muñoz 2023) are common or frequent in Algeria (e.g. Bryum
species percentages in these European countries range from argenteum, Didymodon insulanus, D. vinealis, Funaria hygro-
20.1% to 24.6%, which coincides very well with that of metrica, Ptychostomum capillare, P. pseudotriquetrum, Syntrichia
Algeria and is within the range of the Maghreb countries. ruralis, Tortula subulata…), which leads to the assumption
Concerning the mosses, the percentage of species in the that the country is still underexplored. Also, species such
European countries (from 75.4% to 79.9%) is very similar as Plasteurynchium meridionale, Pohlia cruda, and Pottiopsis
to the Algerian one and is close to the Maghreb countries. caespitosa, which are rare in Algeria (reported from two to
These proportions are within the expected range given that six localities), are expected to be much more abundant, as
the Mediterranean climatic conditions limit the development they are typical Mediterranean species (Dierßen 2001). On
of liverworts, which require a higher level of humidity than the contrary, it’s plausible that certain species reported from
mosses (Frahm 2010). very few localities may stem from misidentifications, cast-
This study shows that Algeria has the lowest species/km2 ing doubt on their reliability regarding their presence in the
ratio (0.20 × 10-3), compared to the rest of the Maghreb and country. Among these species, some haven’t been officially
the selected European Mediterranean countries. It is not accepted and are thus labeled with an asterisk (*) indicating
surprising, as Algeria is the biggest African country. With their doubtful status in Algeria. A critical step would be to
a surface area of 2 381 741 km2, it is 5.3 times larger than review the collections to ascertain their reliability. For recent
Morocco and 14.5 times larger than Tunisia. Algeria is domi- reports like Brachymenium commutatum and Hydrogonium
nated by the Saharan Region, with very harsh weather con- arcuatum, this should be feasible, as they are expected to be
ditions that hinder the development of bryophytes (Ozenda preserved in an herbarium. For older reports, locating speci-
1977). This ratio in the European countries is much higher mens is highly improbable or even impossible, as they may
than in the Maghreb. When looking at the species/km2 ratio be challenging to find or possibly destroyed. Consequently,
in the European Mediterranean countries, even in Greece, these reports can only be reconsidered for inclusion in the
the smallest country considered (110 496 km2), it is much Algerian flora if new collections are discovered. That is the

108 CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5)

 Revisiting the Bryophyte Checklist of Algeria

Table 2. — Dominant families and genera of bryophytes in Algeria.

Number of taxa Number of taxa

Liverworts (% of hornworts and liverworts’ taxa) Mosses (% of mosses’ taxa)
Families
Number of families 32 Number of families 46
Ricciaceae 31 (25.6%) Pottiaceae 107 (28.4%)
Riellaceae 8 (6.6%) Brachytheciaceae 39 (10.3)
Fossombroniaceae 7 (5.8%) Bryaceae 27 (7.2%)
Lophocoleaceae 6 (5.0%) Grimmiaceae 26 (6.9)
Scapaniaceae 6 (5.0%) Orthotrichaceae 24 (6.4%)
Cephaloziellaceae 5 (4.1%) Fissidentaceae 20 (5.3%)
Porellaceae 5 (4.1%) Mniaceae 16 (4.2%)
Funariaceae 15 (4.0%)
Bartramiaceae 10 (2.7%)
Subtotal 68 (56.2%) Subtotal 288 (74.4%)
Other families (25) 50 (43.8%) Other families (37) 93 (24.7%)
Genera
Number of genera 55 Number of genera 135
Riccia 30 (24.8%) Tortula 22 (5.8%)
Riella 8 (6.6%) Fissidens 20 (5.3%)
Fossombronia 7 (5.8%) Ptychostomum 17 (4.5%)
Cephaloziella 5 (4.1%) Grimmia 16 (4.2%)
Porella 5 (4.1%) Orthotrichum 13 (3.4%)
Syntrichia 12 (3.2%)
Weissia 11 (2.9%)
Didymodon 10 (2.7%)
Entosthodon 10 (2.7%)
Subtotal 55 (45.5%) Subtotal 131 (34.7%)
Other genera (50) 50 (54.5%) Other genera (126) 50 (65.3%)

case of Physcomitrium pyriforme, as it is a rare species in the the northern regions of Kabylia and Kroumerie, primarily
Mediterranean and frequently confused with Entosthodon in the Tell Atlas. Among the better-studied mountain ranges
fascicularis (Brugués & Ruiz 2010). Also, the presence of are the Tlemcen Mountains, the Blida Atlas, the Djurdjura
some pleurocarpous mosses seems to be unreliable. That is Massif, the Constantine Mountains, and the Aurès Massif.
the case of Hamatocaulis vernicosus, reported from Gorges To a lesser extent, samples come from the Hauts Plateaux and
de la Chiffa. Despite being a Circumpolar Boreal-montane from locations already included in the Saharan region. How-
species, its ecological preferences for inhabiting mires, flushes, ever, the Hoggar and Tassili n’Ajjer massifs have also attracted
fenny hollows, and springheads in upland areas (Blockeel et al. attention and have been the subject of interesting collections
2014b), do not seem to correspond to the characteristics of and findings, such as Brachymenium exile, Fabronia leikipiae
the Algerian locality. The presence of three Hylocomiaceae var. ahaggarensis, Funaria altissima, Riccia frostii, Vesicularia
reported from Tizi-Ouzou or the neighboring areas in the reimersiana, and Weissia armata.
Kabylie du Djurdjura, such as Hylocomiadelphus triquetrus,
Hylocomium splendens, and Loeskeobryum brevirostre, is also
improbable in localities close to the Mediterranean coast. It CONCLUSIONS
can be hypothesized that because they share a similar habitus,
they could even belong to a unique species. Five Plagiothe- The Algerian bryophyte flora, at present consisting of
ciaceae have been reported in the Kabylie du Djurdjura, a 498 taxa, was made mainly by French botanists in the 19th
very uncommon family in the Mediterranean Region. Two and first half of the 20th centuries. It cannot be consid-
of their species, Isopterygiopsis pulchella and Orthothecium ered well known. The presence of great mountain systems,
intricatum, are likely to be found in Mediterranean areas which include a great diversity of forests in the northern
(Guerra et al. 2018a, b), and therefore they have been coastal region of Algeria, as well as steppe areas with many
accepted but are reported in the same locality (Djurdjura, Ifri temporary lagoons and even desertic areas in the Saharan
n’Ouadnefel). Since they are often confused (Blockeel et al. region, portend a higher diversity of bryophytes than has
2014b; Guerra et al. 2018a), their reliability is uncertain. been reported until now, mainly of well-adapted species,
Also, the presence of Platygyrium repens and Pylaisia poly- as in many other well-known European Mediterranean
antha is doubtful. Both are Circumpolar Temperate species countries. More intensive exploration of the country is
(Blockeel et al. 2014b), and their identification is difficult necessary to accurately assess the true abundance of spe-
to establish without doubt, even to distinguish them from cies, in addition to completing the species diversity. This
some Hypnaceae species. includes confirmation of the presence of certain dubious
From the collection data, it is observed that the most sam- taxa, some of which have only been reported once and
pled areas are the wettest and richest in bryophytes, mainly whose citations cannot be reviewed because herbarium

CRYPTOGAMIE, BRYOLOGIE • 2024 • 45 (5) 109

 Ros R. M. et al.

Table 3. — Comparison in species richness in the Maghreb countries (in the restricted sense) and three European Mediterranean countries.

Maghreb Europe
Algeria Morocco Tunisia Greece Italy Spain
Bryophyte taxa number 498 526 305 743 1240 1182
Liverworts and hornworts taxa number (%) 121 (24.3) 111 (21.1) 91 (29.8) 594 (20.1) 305 (24.6) 286 (24.2)
Mosses taxa number (%) 377 (75.7) 415 (78.9) 214 (70.2) 149 (79.9) 935 (75.4) 896 (75.8)
Surface of the country (km2) 2 381 741 446 550 163 610 110 496 251 408 492 899
Bryophyte taxa/km2 0.20 × 10-3 1.18 × 10-3 1.86 × 10-3 6.72 × 10-3 4.93 × 10-3 2.40 × 10-3

Bell D., Long D. G. & Hollingsworth P. 2013. — The use

collections are difficult or impossible to find due to the of DNA barcoding to address major taxonomic problems for rare
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Submitted on 4 October 2023;

accepted on 29 April 2024;
published on 10 October 2024.

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